58 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 253 3. EN/SVL 0.082 ................ rubra EN/SVL 0.082 .................... 4 4. Canthus rostralis sharp, loreal region nearly vertical, postocular fold straight ......... ..................... schlaginhaufeni Canthus rostralis rounded, loreal region sloping, postocular fold curves down abruptly behind ear ................... dentata 5. Advertisement call a single note repeated every several seconds, males possess vocal slits .................... rhododactyla Advertisement call a series <strong>of</strong> rapidly repeated notes, males lack vocal slits ...... similis Liophryne allisoni, new species Figure 32 HOLOTYPE: AMNH A81221 (field no. RZ 7890), collected by R. G. Zweifel on July 22, 1968, in Morobe Province, Papua New Guinea, on the road from Wau to Edie Creek, north slope <strong>of</strong> Mt. Kaindi, elevation 1900 m, about 5 km west and 1 km north <strong>of</strong> Wau. Paratypes (all from Papua New Guinea): Eastern Highlands Prov.: AMNH A66710– 66718, collected by H. M. Van Deusen (Sixth Archbold Expedition), Oct. 14–24, 1959, at Arau, 1400 m, 22 km E, 9 km S Kainantu. Morobe Prov.: AMNH A83080, AMS 111068, collected by H. G. Cogger (Alpha Helix New Guinea Expedition), Sept. 7, 1969, on the Wau–Edie Creek Road, about 9.7 km from Wau; BPBM 6230, collected by J. L. Gressitt, Sept. 4–6, 1974, 20–25 km ESE Kaisenik, 1700–2000 m; BPBM 6358, collected by A. Allison, June 26, 1974, on Mt. Kaindi, 1800 m, 5 km WNW Wau; BPBM 6417, 6418, collected by T. K. Pratt et al., Oct. 17, 1974, on Mt. Missim, about 10 km NE Wau, 1600 m; BPBM 8475, collected by A. Allison, Jan. 23, 1982, on Mt. Missim, about 1500 m; BPBM 8515, collected by A. Allison, Feb. 19, 1982, on Mt. Missim, 1650 m; BPBM 9359, 9361, 9366, collected by A. Allison, Feb. 16, 1982 on Mt. Missim, Poverty Ck., 1500 m; BPBM 9371, 9374–9376, 9379, collected by A. Allison, Feb. 19, 1983, on Mt. Missim, Coldwater Ck., 1650 m; BPBM 9389, 9390, collected by A. Allison, Feb. 12, 1982, on Mt. Missim, Poverty Ck., 1550 m; BPBM 9408, collected by A. Allison, Feb. 14, 1982, on Mt. Missim, Poverty Ck., 1600 m; BPBM 9550, collected by A. Allison, Apr. 4, 1982, on Mt. Missim, Poverty Ck., 2000 m; BPBM 9554, collected by A. Allison, Feb. 16, 1982, on Mt. Missim, 1990 m; BPBM 9628–9632 (9631 C&S), collected by M. Buka, Apr. 20, 1982, on Mt. Missim, 1950 m; BPBM 9702, collected by A. Allison, May 20, 1982, 5 km NNW Wau, 1800 m. ETYMOLOGY: The patronym honors Dr. Allen Allison <strong>of</strong> the Bishop <strong>Museum</strong>, in recognition <strong>of</strong> his collections <strong>of</strong> this species as well as other important contributions to the biology <strong>of</strong> Papua New Guinea. DIAGNOSIS: Differs from other species <strong>of</strong> Liophryne in its small size (SVL 30 mm or less) and in the small size and relative obscurity <strong>of</strong> the ear. L. dentata and L. schlaginhaufeni just enter maturity at the maximum size <strong>of</strong> allisoni, and similis and rhododactyla do so at a still larger size. DESCRIPTION OF HOLOTYPE: Adult male with the following measurements and proportions: SVL 24.6, HW 10.2, TL 12.9, EY 3.0, EN 2.0, IN 3.1, HD 5.8, FT 12.8, third finger disc 0.55, fourth toe disc 1.0, TY 2.0; HW/SVL 0.415, TL/SVL 0.524, EY/ SVL 0.122, EN/SVL 0.081, IN/SVL 0.126, EN/IN 0.645, HD/SVL 0.236, FT/SVL 0.520, FD/SVL 0.022, TD/SVL 0.041. Head narrower than body, snout bluntly pointed seen from above, rounded and slightly protruding in pr<strong>of</strong>ile; loreal region a moderately steep slope, scarcely concave; canthus rostralis rounded; nostrils lateral, barely visible from above, a little closer to tip <strong>of</strong> snout than to eye. Eye moderately large, eyelid about 80% <strong>of</strong> interorbital space; tympanic annulus scarcely visible. Relative lengths <strong>of</strong> fingers 3 4 2 1, first more than half length <strong>of</strong> second, all with grooved terminal discs scarcely or (1st) not broader than penultimate phalanges; subarticular and metacarpal elevations low, rounded. Toes unwebbed, relative lengths 4 3 5 2 1, all with grooved terminal discs, those on third and fourth about 2 widths <strong>of</strong> penultimate phalanges, others progressively narrower; subarticular elevations rounded, slightly prominent; a small, elongate inner metatarsal tubercle, no others. Skin smooth above and below with only a trace <strong>of</strong> a postocular fold. The ground color in preservative is tan with small, vague, darker marks and a pair <strong>of</strong> more prominent, small dark spots in the
2000 ZWEIFEL: PARTITION OF <strong>SPHENOPHRYNE</strong> 59 scapular region. The side <strong>of</strong> the face is dusky, darker below the eye and around the nostril. A narrow, dark brown postocular streak breaks up above the ear and continues in intermittent fashion down to midflank. There are also some dark spots between the ear and front leg. The edge <strong>of</strong> the chin is dusky with lighter spots. On the throat and chest this gives way to pale brown mottling on the paler background. The abdomen and undersides <strong>of</strong> thighs are unmarked, the shank and tarsus mottled, and the sole and back <strong>of</strong> the tarsus nearly uniform dark brown. There is a well-defined, dark brown, triangular subcloacal patch. In life the dorsal surfaces (head, body, and
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