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SPHENOPHRYNE - American Museum of Natural History

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126 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 253<br />

found in disjunct populations at high elevations.<br />

Too little is known <strong>of</strong> the details <strong>of</strong> geographic<br />

distribution <strong>of</strong> species within New<br />

Guinea to support speculation about distribution<br />

patterns and areas <strong>of</strong> endemism. On<br />

a broader scale, however, the diversity <strong>of</strong><br />

microhylids very likely relates to their reproductive<br />

mode that divorces them from<br />

the need to lay their eggs in water. New<br />

Guinea is an extremely mountainous island<br />

Archbold, R., and A. L. Rand<br />

1935. Results <strong>of</strong> the Archbold Expeditions.<br />

No. 7. Summary <strong>of</strong> the 1933–1934<br />

Papuan Expedition. Bull. Am. Mus.<br />

Nat. Hist. 68(8): 527–579.<br />

Archbold, R., A. L. Rand, and L. J. Brass<br />

1942. Results <strong>of</strong> the Archbold Expeditions.<br />

No. 41. Summary <strong>of</strong> the 1938–1939<br />

New Guinea Expedition. Bull. Am.<br />

Mus. Nat. Hist. 79(3): 197–288.<br />

Bickford, D.<br />

1999. To catch a frog. Wild. Conserv. 102(3):<br />

50–54.<br />

Blommers-Schlösser, R.M.A.<br />

1993. Systematic relationships <strong>of</strong> the Mantellinae<br />

Laurent 1946 (Anura Ranoidea).<br />

Ethol. Ecol. Evol. 5: 199–218.<br />

Blum, J. P., and J. I. Menzies<br />

1988. Notes on Xenobatrachus and Xenorhina<br />

(Amphibia: Microhylidae) from<br />

New Guinea with description <strong>of</strong> nine<br />

new species. Alytes 7(4): 125–163.<br />

Bogart, J. P., and C. E. Nelson<br />

1976. Evolutionary implications from karyotypic<br />

analysis <strong>of</strong> frogs <strong>of</strong> the families<br />

Microhylidae and Rhinophrynidae.<br />

Herpetologica 32(2): 199–208.<br />

Boulenger, G. A.<br />

1897. Descriptions <strong>of</strong> new lizards and frogs<br />

from Mount Victoria, Owen Stanley<br />

Range, New Guinea, collected by Mr.<br />

A. S. Anthony. Ann. Mag. Nat. Hist.<br />

(ser. 6) 19: 6–13.<br />

1898. An account <strong>of</strong> the reptiles and batrachians<br />

collected by Dr. L. Loria in British<br />

New Guinea. Ann. Mus. Civ. Stor.<br />

Nat. Genova (ser. 2) 18: 694–710.<br />

1914. An annotated list <strong>of</strong> the batrachians and<br />

reptiles collected by the British Ornithologist’s<br />

Union and the Wollaston<br />

Expedition in Dutch New Guinea.<br />

REFERENCES<br />

with high rainfall. Under such conditions,<br />

upland standing water habitats are not<br />

abundant, and streams tend to be torrential,<br />

but moist terrestrial and arboreal habitats<br />

are everywhere and are utilized by microhylids<br />

from sea level to the highest mountains.<br />

A parallel situation is in the Admiralty<br />

and Solomon islands, where a diversity<br />

<strong>of</strong> platymantine ranid frogs, all with<br />

direct embryonic development, dominate<br />

the faunas.<br />

Trans. Zool. Soc. London 20(5): 247–<br />

274.<br />

Brass, L. J.<br />

1956. Results <strong>of</strong> the Archbold Expeditions.<br />

No. 75. Summary <strong>of</strong> the Fourth Archbold<br />

Expedition to New Guinea (1953).<br />

Bull. Am. Mus. Nat. Hist. 111(2): 77–<br />

152.<br />

1959. Results <strong>of</strong> the Archbold Expeditions.<br />

No. 79. Summary <strong>of</strong> the Fifth Expedition<br />

to New Guinea (1956–1957). Ibid.<br />

118(1): 1–70.<br />

1964. Results <strong>of</strong> the Archbold Expeditions.<br />

No. 86. Summary <strong>of</strong> the Sixth Archbold<br />

Expedition to New Guinea (1959).<br />

Ibid. 127(4): 145–215.<br />

Brongersma, L. D., and G. F. Venema<br />

1962. To the mountains <strong>of</strong> the stars. London:<br />

Hodder and Stroughton, xv 17–318<br />

pp.<br />

Bulmer, R.N.H., and M. J. Tyler<br />

1968. Karam classification <strong>of</strong> frogs. J. Polynesian<br />

Soc. 77: 333–385.<br />

Burt, C. E., and M. D. Burt<br />

1932. Herpetological results <strong>of</strong> the Whitney<br />

South Seas Expedition. VI. Pacific islands<br />

amphibians and reptiles in the<br />

collection <strong>of</strong> the <strong>American</strong> <strong>Museum</strong> <strong>of</strong><br />

<strong>Natural</strong> <strong>History</strong>. Bull. Am. Mus. Nat.<br />

Hist. 63(5): 461–597.<br />

Burton, T. C.<br />

1984. A new character to distinguish the Australian<br />

microhylid genera Cophixalus<br />

and Sphenophryne. J. Herpetol. 18(2):<br />

206–207.<br />

1986. A reassessment <strong>of</strong> the Papuan subfamily<br />

Asterophryinae (Anura: Microhylidae).<br />

Rec. South Australian Mus.<br />

19(19): 405–450.<br />

1990. The New Guinea genus Copiula Méhely¨<br />

(Anura: Microhylidae): a new di-

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