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Comments Disjunct from Scotland? Edge of GB range? Prop (%) of GB population Wales extant (post-1970) Wales hectads (all records) Significant decline? Europe Red List Section 42 Wales Native/alien status Criteria Wales Red List Taxon MLH GB Red List 2011 GB Red List 2005 EN EN M Bryum gemmiparum EN A, B Native S42 >50% 4 2 33 N Northern world edge in Brecknock, where extant at two sites (River Usk at Sennybridge & Fenni-fach; TBDB), lost from 3 others & from Monmouthshire DD DD M Bryum intermedium CR A, B Native >50% 8 1 1 Lost from dune systems in 6 vice-counties; recently recorded only in Flintshire (Ddol; Hill, 1988) and not looked for there subsequently M Bryum klinggraeffii Native VU VU M Bryum knowltonii CR A, B Native S42 >50% 5 1 5 Lost from 4 sites and now extant as a single small patch at Morfa Dyffryn SSSI (Holyoak, 2001b) NT NT M Bryum kunzei Native 9 7 44 N Northern edge in Anglesey/Caernarfon EN VU M Bryum marratii EN A, B Native S42 RT >50% 4 2 13 S Southern edge in Cardiganshire, lost from 3 sites and now extant over 26x3m in Ynyslas carpark and in a 30m strip near Minfordd (Holyoak, 2001b) M Bryum mildeanum EN A, B Native >50% 8 2 12 S Southern edge in Brecknock, lost from Caernarfon since 1892, only seen twice in south Wales since 1965 (Moel Penderyn 2007 & Foel Fawr 2009; SDSB pers. obs.) M Bryum moravicum Native DD NT M Bryum muehlenbeckii EN B Native 1 1 8 S Yes Southern edge in Caernarfon (Rhaedr Ogwen 1988 & Carnedd Llewelyn 1988; TBDB), otherwise no further south than Perthshire M Bryum pallens Native M Bryum pallescens Native M Bryum pseudotriquetrum var. bimum Native 12 8 8 Under-recorded because of identification difficulties M Bryum pseudotriquetrum var. Native pseudotriquetrum M Bryum radiculosum Native M Bryum riparium Native Rare 11 9 16 S Southern edge in Glamorgan (Cwm Dimbath; Perry, 1994) M Bryum rubens Native M Bryum ruderale Arch? M Bryum sauteri Native M Bryum subapiculatum Native M Bryum tenuisetum Native DD 16 15 23 Scattered in north Wales (Hill, 1988) and south to Radnorshire; easily overlooked M Bryum torquescens Native 9 6 15 Rare on carboniferous limestone near the coasts of north and south Wales RE RE M Bryum turbinatum EX Native >50% 5 0 0 Lost from Brecknock, Monmouthshire & Meirionydd, now extinct in GB CR RE M Bryum uliginosum EX Native RT >50% 2 0 0 Lost from Meirionydd (Morfa Dyffryn to 1904; Holyoak, 2001b), almost extinct in GB M Bryum violaceum Arch? VU NT M Bryum warneum EN A, B Native S42 Rare >50% 8 3 15 Extant at 2 SSSI in Meirionydd (Morfa Dyffryn & Morfa Harlech) and 1 in Anglesey (Tywyn Aberffraw), lost from 7 other sites (Holyoak, 2001b) M Bryum weigelii EN B Native 2 2 3 One outlier in Shropshire, otherwise southern edge in Caernarfon (six sites in Carneddau; Hill, 1988) M Buxbaumia aphylla EN A, B Native RT >50% 3 1 3 Lost from Meirionydd (upper Dyfi 1896) and Montgomeryshire (Breidden Hill 1960s), and could not be refound in Radnorshire in 2007 having been seen in 1999 (Burfa Bank; Lawley, 2007) M Calliergon cordifolium Native M Calliergon giganteum Native M Calliergonella cuspidata Native M Calliergonella lindbergii Native L Calypogeia arguta Native L Calypogeia azurea VU- B- Native 7 5 5 S Southern edge in Brecknock (Cefn Cwm Crew; Woods, 2006) L Calypogeia fissa Native L Calypogeia muelleriana Native L Calypogeia neesiana Native L Calypogeia sphagnicola Native M Campyliadelphus chrysophyllus Native M Campyliadelphus elodes Native RT 16 10 11 Still locally abundant in fens on Anglesey (S.D.S. Bosanquet pers. obs.) and in dunes in Carmarthenshire (Pembrey) M Campylium protensum Native M Campylium stellatum Native M Campylophyllum calcareum Native M Campylopus atrovirens var. atrovirens Native M Campylopus brevipilus Native 21 17 6 Widespread but generally rare in coastal heathland; Hill (1988) reports it as "locally abundant near Holyhead and at the tip of Lleyn". M Campylopus flexuosus Native 31
32 Comments Disjunct from Scotland? Edge of GB range? Prop (%) of GB population Wales extant (post-1970) Wales hectads (all records) Significant decline? Europe Red List Section 42 Wales Native/alien status Criteria Wales Red List Taxon MLH GB Red List 2011 GB Red List 2005 M Campylopus fragilis Native M Campylopus gracilis Native 11 9 8 S Southern edge in Cardiganshire (Maesnant, Pumlimon; Hill, 1988), rare in north Wales (Hill, 1988) M Campylopus introflexus NA Neo M Campylopus pilifer Native 10 10 37 Present in six places in Pembrokeshire (Bosanquet, 2010) and several on the Barmouth-Porthmadog coast (Hill, 1988), but intermediates to C. introflexus are confusing M Campylopus pyriformis Native M Campylopus setifolius Native Rare 14 12 17 S Southern edge in Meirionydd, occasional in north Wales (Hill, 1988) M Campylopus subulatus Native 27 15 19 A widespread weed of damp gravelly ground, probably overlooked M Campylostelium saxicola EN A, B Native Rare >50% 8 3 9 Lost from Meirionydd (2 sites to 1911) and sites in Caernarfon (Hill, 1988) and Brecknock (Woods, 2006) probably extant in Brecknock (Talybont & Grwyne Fawr), Cardiganshire (Ponterwyd) & Caernarfon (Moel Hebog & Eryri) M Catoscopium nigritum VU D2 Native 2 2 6 S Southern edge on Anglesey (Tywyn Aberffraw & Newborough Warren; Hill, 1988) L Cephalozia bicuspidata Native L Cephalozia catenulata Native L Cephalozia connivens Native L Cephalozia leucantha EN A, B Native >50% 4 1 1 A few Cornish colonies, otherwise southern edge in Meirionydd (Arthog Bog) L Cephalozia loitlesbergeri VU D2 Native 4 4 8 S Southern edge in Cardiganshire (Cors Caron) L Cephalozia lunulifolia Native L Cephalozia macrostachya var. EN A, B Native >50% 7 3 8 Lost from Pembrokeshire (3 sites), still thriving on Cors Fochno (S.D.S. Bosanquet pers. obs., 2009), apparent decline may be because some macrostachya records not assigned to variety L Cephalozia macrostachya var. spiniflora VU D2 Native 2 2 14 Only Welsh records are from Cardiganshire (Cors Caron 1990) and Anglesey (heath nr Brynrefail 1994) L Cephalozia pleniceps Native 21 19 28 Found to be frequent in south Wales suggesting it is overlooked elsewhere (Bosanquet & Motley, 2005), unlikely to be threatened VU NT L Cephaloziella calyculata VU D2 Native S42 Rare 3 3 25 N Northern world edge in Pembrokeshire (2 sites), also in Glamorgan (2 sites) (Bosanquet, 2008b) L Cephaloziella divaricata Native L Cephaloziella elachista VU D2 Native DD 1 1 8 W Western edge in Pembrokeshire, where extant at only Welsh site (Esgyrn Bottom) L Cephaloziella hampeana Native NT VU L Cephaloziella massalongi CR A, B Arch? Rare >50% 9 1 7 N Northern edge on Anglesey (last seen 1959), still extant at Figra Mine (Des Callaghan pers. obs., 2011) otherwise last seen near Llanfrothen in 1967 (TBDB) VU VU L Cephaloziella nicholsonii EN A, B Arch? S42 Rare >50% 2 1 6 N Lost from Meirionydd (Bontddu) since 1923 (Hill, 1988), now northern world edge in Cardiganshire (Cwmystwyth 1998) L Cephaloziella rubella Native L Cephaloziella spinigera EN A, B Native >50% 2 1 4 Lost from Anglesey (Llandonna Common), perhaps extant at single site in Cardiganshire (Cors Caron 1970; Hale, 1998) L Cephaloziella stellulifera Native 31 22 24 Relatively frequent on the Welsh coast and probably somewhat under-recorded NT NT L Cephaloziella turneri NT B Native 11 6 23 One outlier on Kintyre, otherwise northern edge in Carmarthenshire (Laugharne), lost from Meirionydd (Y Gardd, Minffordd) since 1916 (Hill, 1988) and from sites in Carmarthenshire & Monmouthshire M Ceratodon purpureus Native L Chiloscyphus pallescens Native L Chiloscyphus polyanthos Native M Cinclidium stygium RE Native >50% 1 0 0 Lost from Meirionydd (Cadair Idris 1939; Hill, 1988) M Cinclidotus fontinaloides Native M Cirriphyllum crassinervium Native M Cirriphyllum piliferum Native L Cladopodiella fluitans Native L Cladopodiella francisci EN A, B Native >50% 7 3 4 Lost from Monmouthshire, Meirionydd & 1 site in Pembrokeshire, extant at single sites in Glamorgan, Pembrokeshire & Anglesey M Climacium dendroides Native L Cololejeunea calcarea Native 28 18 7 S Southern edge in Monmouthshire L Cololejeunea minutissima Native L Cololejeunea rossettiana Native 10 9 13 Reasonably frequent on shaded carboniferous limestone L Colura calyptrifolia Native RT 48 43 24 Dramatic recent increase, especially in conifer plantations L Conocephalum conicum Native
Page 1 and 2: 1 Plantlife Cymru Uned 14, Llys Cas
Page 3 and 4: A Bryophyte Red Data List for Wales
Page 31: 30 Comments Disjunct from Scotland?
Page 35 and 36: 34 Comments Disjunct from Scotland?

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