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3.2 Alien status<br />
Only six Welsh bryophyte species were considered to be of recent introduced origin by Hill<br />
et al. (2007) (i.e. they are neophytes, meaning they were introduced after 1500AD):<br />
Atrichum crispum, Campylopus introflexus, Didymodon umbrosus, Hennediella<br />
stanfordensis, Lophocolea semiteres and Orthodontium lineare. The Didymodon and<br />
Lophocolea are rare in Wales but have not been assigned to a threat category because of<br />
their non-native status. It is possible that other Welsh bryophytes are archaeophytes<br />
(introduced before 1500AD), especially some arable taxa and metalophytes (species of<br />
substrates rich in heavy metals, such as some mine spoil). Hill et al. considered<br />
Anthoceros agrestis, Bryum ruderale, Bryum violaceum, Cephaloziella massalongi*,<br />
Cephaloziella nicholsonii*, Dicranella staphylina, Didymodon tomaculosus*, Ditrichum<br />
plumbicola, Phaeoceros carolinianus*, Scopelophila cataractae*, Targionia hypophylla<br />
and Weissia squarrosa* to be possible archaeophytes, but the lack of proof means that all<br />
have been included in the Red Data List analysis and several of them (marked *) have<br />
been assigned a threat category.<br />
3.3 Geographic coverage<br />
This Red Data List covers the country of Wales, including the 13 vice-counties of<br />
Monmouthshire (vice-county number 35), Glamorgan (41), Brecknockshire (42),<br />
Radnorshire (43), Carmarthenshire (44), Pembrokeshire (45), Cardiganshire (46),<br />
Montgomeryshire (47), Merionethshire (48), Caernarvonshire (49), Denbighshire (50),<br />
Flintshire (51) and Anglesey (52).<br />
In the absence of more detailed tetrad recording, decline statistics were based on data for<br />
Welsh hectad (10 km square of the Ordnance Survey grid) records from the British<br />
Bryological Society database held by the Biological Records Centre. Almost all these<br />
records are correctly attributed to a vice-county, but a few records from the Welsh border<br />
result from hectad record cards put together for the bryophyte Atlas (Hill et al., 1991-<br />
1994) and could come from England (R.G. Woods, pers. comm.). The western distribution<br />
of most of Wales’ threatened bryophytes means that <strong>this</strong> is unlikely to be much of an<br />
issue when assessing declines.<br />
Rhestr Data Coch Bryoffytau ar gyfer Cymru<br />
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