INTRODUCTION Vegetation, land use and rural livelihoods The rainfall and temperature variability, in relation to the rapid rate <strong>of</strong> population growth in East Africa (~7.1% per annum), significantly determines the livelihood strategies <strong>of</strong> indigenous food producers (e.g., cultivators, livestock keepers and fishers) who exploit the country’s natural resources (Conway et al., 2005; Thornton et al., 2008) (Fig. iii). In total, about 70 to 80% <strong>of</strong> the East African workforce is employed in the agricultural sector (FAOSTAT, 2009), involving many distinct crop-livestock systems exploiting a wide range <strong>of</strong> agro-products (e.g., banana, maize, sorghum, millet, potatoes, meat, milk). Over 75% <strong>of</strong> total agricultural output is provided by smallholder farmers, with farm sizes <strong>of</strong> about 2.5 ha on average, mainly producing for home-consumption, and to a lesser degree for (semi-)commercial sale <strong>of</strong> goods. The farming activities are completely organised and directed within the family, operating within a network <strong>of</strong> relations at the community level and using limited technological innovations (Salami et al., 2010). Intensive small-scale subsistence farming is the most widespread permanent agricultural system (Widgren and Sutton, 2004) found in areas with high population density (Okigbo, 1990), such as the Naya in northwestern Tanzania (Maruo, 2002), the Chagga in north Tanzania (Fernandes et al., 1984) and the Ankole in southwestern Uganda (Kasfir, 1993). In sparsely populated semi-arid regions, such as the Ngorongoro district in north Tanzania and the Narok district in southwestern Kenya, cattle breeding forms an essential part <strong>of</strong> the rural livelihoods (Dixon et al., 2001). To optimise the use <strong>of</strong> rangeland resources for meat and milk production, transhumant pastoralists, such as the Maasai, move their herds throughout the year, maintaining semi-permanent home bases. In times <strong>of</strong> drought and during the night, livestock enclosures protect the herds from predators and from raiding by other humans (Marshall, 2000; Butt, 2010). In the last few decades, many <strong>of</strong> East Africa’s life-support have experienced dynamic changes induced by environmental, socioeconomic and political impacts (Maruo, 2002). Climate variability, however, is likely to have the most severe impact on resource-poor agriculturalists, leading to changes in productivity <strong>of</strong> rainfed crops and forage, reduced water availibility and severity <strong>of</strong> crop, livestock and human diseases (Goulden, 2005; Thornton et al., 2008). Moreover, soil erosion has severely increased as a result <strong>of</strong> land-use intensification (over-cultivation, over-grazing, logging), land fragmentation (loss <strong>of</strong> buffer zones) and the lack <strong>of</strong> fertilisers (Pomeroy et al., 2003; Ngecu et al., 2004), and is detrimental for soil nutrients and food production potential (Toy et al., 2002). Land degradation and the accelerating conversion <strong>of</strong> natural lands for agricultural purposes also causes a substantial loss <strong>of</strong> plant, mammal and bird diversity (Pomeroy et al., 2003). The threat <strong>of</strong> extinction may be particularly acute for the more than a million estimated species <strong>of</strong> fungi living in direct association with plants (e.g., Hawksworth, 1991, 1993; Hyde and Hawksworth, 1997). For example, based on limited data in Uganda the rate <strong>of</strong> overall biodiversity loss is estimated to have reached 10% per decade, with even higher values in savanna systems (~20%) and agro-ecosystems (~50%) (Pomeroy and Mwima, 2002). Also surface water resources, such as lakes and wetlands, become progressively depleted by eutrophication and/or siltation due to the combined impact <strong>of</strong> human activities and climate variability (e.g., Verschuren et al., 2002; Ngaira, 2009; Stager et al., 2009). Anomalies in the nutrient cycling <strong>of</strong> large water bodies, such as L. Tanganyika, severely affect fish abundance and fishery (O’Reilly et al., 2003). A recent more qualitative and quantitative assessment <strong>of</strong> the biogeochemical cycle in L. Tanganyika revealed that anthropogenic global warming may possibly be the main driver <strong>of</strong> decreasing lake productivity over the past 90 years (Tierney et al., 2010). Thus, by studying current and past ecosystem responses to human impact and climate change, strategies necessary for successful community-based natural resource management can be developed to retain resilience <strong>of</strong> the ecosystems, which support them (e.g., Willis and Bhagwat, 2010). . 22 .
1 Figure 2 Global Land-Cover 2000 map <strong>of</strong> Africa. Fig. iii. Vegetation map <strong>of</strong> Africa, with special reference to the East african region, which is mainly characterised by cropland, mosaic forest, deciduous woodland/shrubland and open grassland (Mayaux et al., 2004). . 23 . New land-cover map <strong>of</strong> Africa Journal <strong>of</strong> Biogeography 31, 861–877, ª 2004 Blackwell Publishing Ltd 869 INTRODUCTION
- Page 3: 2011 Thesis submitted in partial fu
- Page 7: Voorwoord . 7 . VOORWOORD Aarde, wa
- Page 10: DANKWOORD / ACKNOWLEDGEMENTS door K
- Page 14 and 15: TABLE OF CONTENTS A.1. Septated spo
- Page 17 and 18: Abstract . 17 . ABSTRACT In tropica
- Page 19 and 20: Introduction . 19 . INTRODUCTION
- Page 21: . 21 . INTRODUCTION Atlantic Ocean
- Page 25 and 26: Fig. iv. Rainfall variation in east
- Page 27 and 28: . 27 . INTRODUCTION metallurgy main
- Page 29 and 30: . 29 . INTRODUCTION analysis of fos
- Page 31 and 32: . 31 . INTRODUCTION (iv) illuminate
- Page 33 and 34: . 33 . INTRODUCTION Coniochaeta spe
- Page 35 and 36: . 35 . INTRODUCTION university pres
- Page 37 and 38: . 37 . INTRODUCTION Mumbi, C.T., Ma
- Page 39 and 40: . 39 . INTRODUCTION lipids in an eq
- Page 41 and 42: . 41 . CHAPTER 1 Diversity and ecol
- Page 43 and 44: Equator b 0° Altitude (m) a >3000
- Page 45 and 46: . 45 . CHAPTER 1 Diversity and ecol
- Page 47 and 48: 1.3.1. Descriptions and illustratio
- Page 49 and 50: Type HdV-1005: Brachydesmiella sp.
- Page 51 and 52: Type HdV-1020 (Plate II) . 51 . CHA
- Page 53 and 54: . 53 . CHAPTER 1 Diversity and ecol
- Page 55 and 56: Type HdV-1051 (Plate III) . 55 . CH
- Page 57 and 58: . 57 . CHAPTER 1 Diversity and ecol
- Page 59 and 60: . 59 . CHAPTER 1 Diversity and ecol
- Page 61 and 62: . 61 . CHAPTER 1 Diversity and ecol
- Page 63 and 64: RefeRences . 63 . CHAPTER 1 Diversi
- Page 65 and 66: . 65 . CHAPTER 1 Diversity and ecol
- Page 67 and 68: . 67 . CHAPTER 1 Diversity and ecol
- Page 69: . 69 . CHAPTER 1 Diversity and ecol
- Page 72 and 73:
CHAPTER 2 Modern non-pollen palynom
- Page 74 and 75:
CHAPTER 2 Modern non-pollen palynom
- Page 76 and 77:
CHAPTER 2 Modern non-pollen palynom
- Page 78 and 79:
CHAPTER 2 Modern non-pollen palynom
- Page 80 and 81:
CHAPTER 2 Modern non-pollen palynom
- Page 82 and 83:
CHAPTER 2 Modern non-pollen palynom
- Page 84 and 85:
CHAPTER 2 Modern non-pollen palynom
- Page 86 and 87:
CHAPTER 2 Modern non-pollen palynom
- Page 88 and 89:
CHAPTER 2 Modern non-pollen palynom
- Page 90 and 91:
CHAPTER 2 Modern non-pollen palynom
- Page 92 and 93:
CHAPTER 2 Modern non-pollen palynom
- Page 94 and 95:
CHAPTER 2 Modern non-pollen palynom
- Page 96 and 97:
CHAPTER 2 Modern non-pollen palynom
- Page 98 and 99:
CHAPTER 2 Modern non-pollen palynom
- Page 100 and 101:
CHAPTER 2 Modern non-pollen palynom
- Page 102 and 103:
CHAPTER 2 Modern non-pollen palynom
- Page 104 and 105:
CHAPTER 2 Modern non-pollen palynom
- Page 106 and 107:
CHAPTER 2 Modern non-pollen palynom
- Page 108 and 109:
CHAPTER 2 Modern non-pollen palynom
- Page 110 and 111:
CHAPTER 2 Modern non-pollen palynom
- Page 112 and 113:
CHAPTER 2 Modern non-pollen palynom
- Page 114 and 115:
CHAPTER 2 Modern non-pollen palynom
- Page 116 and 117:
CHAPTER 2 Modern non-pollen palynom
- Page 118 and 119:
CHAPTER 2 Modern non-pollen palynom
- Page 120 and 121:
CHAPTER 2 Modern non-pollen palynom
- Page 122 and 123:
CHAPTER 2 Modern non-pollen palynom
- Page 124 and 125:
CHAPTER 2 Modern non-pollen palynom
- Page 126 and 127:
CHAPTER 2 Modern non-pollen palynom
- Page 128 and 129:
CHAPTER 3 Effects of Land use on th
- Page 130 and 131:
CHAPTER 3 Effects of Land use on th
- Page 132 and 133:
CHAPTER 3 Effects of Land use on th
- Page 134 and 135:
CHAPTER 3 Effects of Land use on th
- Page 136 and 137:
CHAPTER 3 Effects of Land use on th
- Page 138 and 139:
CHAPTER 3 Effects of Land use on th
- Page 140 and 141:
CHAPTER 3 Effects of Land use on th
- Page 142 and 143:
CHAPTER 3 Effects of Land use on th
- Page 144 and 145:
CHAPTER 3 Effects of Land use on th
- Page 146 and 147:
CHAPTER 3 Effects of Land use on th
- Page 148 and 149:
CHAPTER 3 Effects of Land use on th
- Page 150 and 151:
CHAPTER 3 Effects of Land use on th
- Page 152 and 153:
CHAPTER 3 Effects of Land use on th
- Page 154 and 155:
CHAPTER 3 Effects of Land use on th
- Page 156 and 157:
CHAPTER 3 Effects of Land use on th
- Page 158 and 159:
CHAPTER 3 Effects of Land use on th
- Page 160 and 161:
CHAPTER 3 Effects of Land use on th
- Page 162 and 163:
CHAPTER 4 Validation of non-pollen
- Page 164 and 165:
CHAPTER 4 Validation of non-pollen
- Page 166 and 167:
CHAPTER 4 Validation of non-pollen
- Page 168 and 169:
CHAPTER 4 Validation of non-pollen
- Page 170 and 171:
CHAPTER 4 Validation of non-pollen
- Page 172 and 173:
CHAPTER 4 Validation of non-pollen
- Page 174 and 175:
CHAPTER 4 Validation of non-pollen
- Page 176 and 177:
CHAPTER 4 Validation of non-pollen
- Page 178 and 179:
CHAPTER 4 Validation of non-pollen
- Page 180 and 181:
CHAPTER 4 Validation of non-pollen
- Page 182 and 183:
CHAPTER 4 Validation of non-pollen
- Page 184 and 185:
CHAPTER 4 Validation of non-pollen
- Page 187 and 188:
General discussion and conclusion .
- Page 189 and 190:
. 189 . GENERAL DISCUSSION AND CONC
- Page 191 and 192:
. 191 . GENERAL DISCUSSION AND CONC
- Page 193 and 194:
. 193 . GENERAL DISCUSSION AND CONC
- Page 195:
. 195 . GENERAL DISCUSSION AND CONC
- Page 198 and 199:
SAMENVATTING beperkingen (i.v.m. de
- Page 200 and 201:
APPENDICES Appendix I. Raw NPP data
- Page 202 and 203:
APPENDICES Mahuhura Kitere Katanda
- Page 204 and 205:
APPENDICES Mahuhura Kitere Katanda
- Page 206 and 207:
APPENDICES Mahuhura Kitere Katanda
- Page 208 and 209:
APPENDICES Mahuhura Kitere Katanda
- Page 210 and 211:
APPENDICES Mahuhura Kitere Katanda
- Page 212 and 213:
APPENDICES Appendix II. Raw NPP dat
- Page 214 and 215:
APPENDICES L. Chibwera Identificati
- Page 216 and 217:
APPENDICES L. Chibwera Identificati
- Page 218 and 219:
APPENDICES Appendix III. Raw NPP da
- Page 220 and 221:
APPENDICES L. Kanymukali Identifica
- Page 222 and 223:
APPENDICES L. Kanymukali Identifica
- Page 224:
Photograph by Bob Rumes Front cover