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REGIONAL MEETINGS - Natural History Museum

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other calcifuges such as Blechnum spicant, Athyrium filix-femina, Dryopteris dilatata, D. filixmas<br />

and Oreopteris limbosperma, there were also, due to the mixed geology of Arran,<br />

ferns that prefer more basic conditions – Asplenium adiantum-nigrum, A. scolopendrium,<br />

A. trichomanes subsp. quadrivalens, Phegopteris connectilis, Polypodium vulgare, and<br />

Polystichum setiferum, which seemed so out of place among the rocks of the raised beach that<br />

I am ashamed to admit I did not recognise it. More interestingly we found Dryopteris aemula<br />

with its distinctly crisped fronds and both filmies (Hymenophyllum tunbrigense and<br />

H. wilsonii) and of course Mary and others soon found Trichomanes speciosum gametophytes.<br />

The sporophyte is also present on Arran at four isolated sites but understandably their<br />

whereabouts are kept secret, even from such distinguished pteridologists as ourselves.<br />

So far I have not mentioned Dryopteris affinis. Arran has all three subspecies and numerous<br />

hybrids and variants, and with Ken Trewren as one of our party we were not going to get<br />

away with recording the species alone. So as well as D. affinis subspp. affinis and borreri,<br />

we had subsp. cambrensis var. paleaceocrispa (the normal cambrensis according to Ken),<br />

and (later confirmed by Ken) subsp. cambrensis var. pseudocomplexa (otherwise known as<br />

morphotype arranensis of Pigott).<br />

Further evidence of the complexity of the Dryopteris affinis agg. on Arran was found in<br />

Cordon Wood (26/035296) in the afternoon. Here, led by the indefatigable Tony Smith up<br />

and down the steep slopes of this seemingly impenetrable woodland, we added D. affinis subsp.<br />

affinis var. paleaceolobata and D. x complexa nothosubsp. complexa (along with its parents) as<br />

well as the D. affinis subspecies and variants we had seen in the morning. By this time not a few<br />

of us were becoming confused, and please, if you wish further elucidation, consult Ken, not me.<br />

Cordon Wood also revealed some more easily recognisable ferns such as Dryopteris aemula<br />

again, and of course D. dilatata, Athyrium filix-femina, Blechnum spicant, Oreopteris<br />

limbosperma and Pteridium aquilinum. We also noted Asplenium trichomanes subsp.<br />

quadrivalens, Hymenophyllum tunbrigense, Phegopteris connectilis and Polypodium<br />

interjectum, while Martin and Mary found some fine examples of Polystichum aculeatum<br />

that the rest of us inexplicably missed. Ken, who had sensibly brought a torch, searched for<br />

and found the Trichomanes gametophyte again. Cordon Wood was the only place,<br />

surprisingly, where we recorded horsetails, though they surely must be abundant on Arran.<br />

Here we saw Equisetum arvense and E. sylvaticum.<br />

316<br />

photo: F. McGavigan<br />

Glen Sannox, Arran<br />

Tony Smith, Martin Rickard, Mary Gibby, Christine Nicholson, Ken Trewren,<br />

Frank Katzer, Alastair Wardlaw, Roland Ennos, Liz Doig, Angus Hannah

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