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Maps produced by kriging and other suitable interpolation methods were essentially similar. Spatial autocorrelation analysis was carried out using the Spatial Autocorrelation Analysis Program (SAAP) version 4.3 (Wartenberg, 1989). This package produces correlograms of autocorrelation coefficient values against each distance class. Moran's `i' and Geary's 'c' autocorrelation coefficients (Cliff and Ord, 1973) were calculated in the present study since these are the most commonly used and have been recommended for use in ecological applications (Sokal, 1979; Hubert et al., 1981; Legendre and Fortin, 1989). The SAAP program gives i values in the range 1 to -1 and c values in the range 0-2; while low values of i correspond to negative autocorrelation and high values correspond to positive autocorrelation, the opposite applies for values of c. Each coefficient is sensitive to slightly different behaviours of the data (Jumars et al., 1977). For example, i bears a close resemblance to a Pearson product moment correlation coefficient and is therefore most sensitive to extreme values in the data set while c, which is a distance-type function, is most sensitive to the proximity of similar or dissimilar values, regardless of their departure from the mean (Jumars eta!., 1977). Following Oden (1984) the overall significance of each spatial autocorrelogram was assessed by checking that the most significant spatial autocorrelation coefficient found in a correlogram was significant at a Bonferroni-corrected significance level a', where a'=a/k, k being the number of autocorrelation coefficients (i.e., the number of distance classes) in the correlogram (Sokal, 1986). Spatial autocorrelation analysis allowed the nature of patterns and estimates of patch sizes to be made in this study. The spatial characteristics of patches indicated by autocorrelation analysis were only accepted if the contour plots were consistent with these patterns. 26

RESULTS Scales of Spatial Variability - Transect Survey From the 48 faunal samples taken for this survey, 3062 individuals were collected from a total of 48 taxa. By far the most abundant (2508 individuals) and diverse group was the polychaetes, 26 species from 13 families. The second most abundant group was the molluscs (262 individuals) with 8 species from 4 families. 257 individuals were collected from the crustaceans; 12 species from 7 families. Most of the species sampled were too rare for numerical analyses, i.e., means of

Maps produced by kriging and other suitable interpolation methods were essentially<br />

similar.<br />

Spatial autocorrelation analysis was carried out using the Spatial Autocorrelation<br />

Analysis Program (SAAP) version 4.3 (Wartenberg, 1989). This package produces<br />

correlograms of autocorrelation coefficient values against each distance class.<br />

Moran's `i' and Geary's 'c' autocorrelation coefficients (Cliff and Ord, 1973) were<br />

calculated in the present study since these are the most commonly used and have been<br />

recommended for use in ecological applications (Sokal, 1979; Hubert et al., 1981;<br />

Legendre and Fortin, 1989). The SAAP program gives i values in the range 1 to -1<br />

and c values in the range 0-2; while low values of i correspond to negative<br />

autocorrelation and high values correspond to positive autocorrelation, the opposite<br />

applies for values of c. Each coefficient is sensitive to slightly different behaviours of<br />

the data (Jumars et al., 1977). For example, i bears a close resemblance to a Pearson<br />

product moment correlation coefficient and is therefore most sensitive to extreme<br />

values in the data set while c, which is a distance-type function, is most sensitive to<br />

the proximity of similar or dissimilar values, regardless of their departure from the<br />

mean (Jumars eta!., 1977).<br />

Following Oden (1984) the overall significance of each spatial autocorrelogram was<br />

assessed by checking that the most significant spatial autocorrelation coefficient found<br />

in a correlogram was significant at a Bonferroni-corrected significance level a', where<br />

a'=a/k, k being the number of autocorrelation coefficients (i.e., the number of<br />

distance classes) in the correlogram (Sokal, 1986). Spatial autocorrelation analysis<br />

allowed the nature of patterns and estimates of patch sizes to be made in this study.<br />

The spatial characteristics of patches indicated by autocorrelation analysis were only<br />

accepted if the contour plots were consistent with these patterns.<br />

26

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