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polychaetes on the initial successional dynamics and colonisation mode of the most<br />

abundant invertebrate taxa are addressed.<br />

Chapter 7 considers the micro-scale spatial heterogeneity of P. elegans. Specifically,<br />

the spatial distribution of P. elegans within small-scale patches was monitored<br />

through the year and the importance of adult-juvenile interactions, interspecific<br />

interactions and abiotic variables in determining micro-scale patterns are discussed.<br />

In Chapter 8, the ecological significance of the small-scale P. elegans patches were<br />

investigated. In particular, the structures of patch and non-patch communities were<br />

compared using univariate and multivariate analyses using data from samples taken at<br />

various times during this study. In this chapter, the community structure during the<br />

decline of these patches are studied, giving an insight into the possible causes of their<br />

demise.<br />

Chapter 9 is a general discussion in which the findings of all the preceding chapters<br />

are brought together.<br />

PYGOSPIO ELEGANS<br />

Pygospio elegans Claparede is a small, sedentary, tube-building polychaete. This<br />

species is particularly interesting to study since various studies have suggested that it<br />

possesses a wide habitat tolerance, a variety of feeding mechanisms and displays a<br />

remarkable diversity of reproductive strategies (Morgan, 1997), in addition to its<br />

ability to form patches of increased density, or tube-beds.<br />

Pygospio elegans has been found in a wide range of habitats from vertical rock<br />

crevices (Gudmundsson, 1985) to hard sand and gravel, but reaches its highest<br />

densities in sandy or mixed sediments (Smidt, 1951; Muus, 1967). The species is<br />

common in both marine and brackish waters although the highest densities are found<br />

in the latter, being abundant at salinities as low as 2ppt (Hempel, 1957).<br />

8

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