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succession of annual <strong>crop</strong>s, three inter<strong>crop</strong> management options were compared, namely i)<br />

conventionally tilled inter<strong>crop</strong>s <strong>with</strong> bare soil in winter, ii) tilled inter<strong>crop</strong>s <strong>with</strong> a cover <strong>crop</strong><br />

in winter (used to reduce nitrogen leaching) that may also change the weed growth conditions<br />

(Liebman and Davis, 2000; Moonen and Barberi, 2004) and iii) untilled overwinter stubble<br />

fields (OSFs) corresponding to an agri-environment scheme where the soil is only tilled at the<br />

end of winter (Critchley et al., 2004; Marsall et al., 2007). In such OSFs, established weed<br />

plants may benefit from the absence of soil tillage (as in perennial <strong>crop</strong>s) but also from the<br />

limited inter-plant competition, which may increase their seed output. This agri-environment<br />

scheme may be favourable to farmland biodiversity, as plant residues and seeds remain at the<br />

soil surface where it may be eaten by animals (Moorcroft et al., 2002; Orlowski, 2006).<br />

This experimental <strong>de</strong>sign was used to analyse the temporal dynamics of the emerged weed<br />

communities during the whole experimental period, concentrating on weed species<br />

composition, weed plant <strong>de</strong>nsities, and biomass. Finally, the potential un<strong>de</strong>rlying mechanisms<br />

are discussed, based on the weed population dynamics in the different experimental<br />

treatments.<br />

C.II.2 Methods<br />

C.II.2.1 Experimental <strong>de</strong>sign<br />

The field experiment was located at the experimental farm ‘Epoisses’ of INRA-Dijon in<br />

eastern France (47°20’N, 5°20’E) <strong>with</strong> a semi-continental climate and a calcareous clayey<br />

soil. Nine <strong>crop</strong> treatments were compared (T2-T11, see Table 6 for <strong>de</strong>tails). Treatments varied<br />

first by the <strong>crop</strong> type, opposing a succession of annual <strong>crop</strong>s: winter wheat (Triticum<br />

aestivum)–spring barley (Hor<strong>de</strong>um vulgare)–summer soybean (Glycine max) and two PFCs:<br />

alfalfa/lucerne (Medicago sativa) and cocksfoot/orchard grass (Dactylis glomerata).<br />

Within<br />

the perennial <strong>crop</strong>s, treatments further varied by <strong>crop</strong> sowing season, opposing autumn sowing<br />

(4 Sept. 2006) and spring sowing (27 April 2007), and by cutting frequency, opposing a high<br />

frequency (5 cuttings per year, C+) and a low frequency (3 cuttings per year, C-) for the<br />

autumn sown plots. Sowing and cutting dates are given in Table 6,<br />

lower part. Cuttings were<br />

performed at about 3-8cm height from soil surface using a forage mower adapted to the small<br />

experimental plots that directly removed the cut biomass. Within the succession of annual<br />

<strong>crop</strong>s, three inter<strong>crop</strong> treatments were compared: treatment T9 <strong>with</strong> superficial soil tillage (5-<br />

8cm) performed <strong>with</strong> a rotary hoe ‘rotavator’ after <strong>crop</strong> harvest (‘conventional’, bare soil<br />

73

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