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Diversifying crop rotations with temporary grasslands - Université de ...

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forage <strong>crop</strong>s sometimes cited the increased competition or the mowing or grazing activities as<br />

possible causes (e.g., Norris and Ayres, 1991; Schoofs and Entz, 2000), while increased<br />

abundances were sometimes linked to reduced herbici<strong>de</strong> use or reduced soil tillage (e.g.,<br />

Cardina et al., 2002a; Bellin<strong>de</strong>r et al., 2004). Few <strong>de</strong>tails were given for i<strong>de</strong>ntifying which<br />

phases of the weed life cycle were mostly affected by PFCs. Only Heggenstaller and Liebman<br />

(2006) showed that alfalfa reduced the seedling survival and fecundity of Abutilon<br />

theophrasti, the most important weed species in their system.<br />

A.IV.2 Hypothetical mechanisms causing the impacts<br />

Annual and perennial <strong>crop</strong>s differ in several important aspects concerning both the<br />

characteristics of the <strong>crop</strong> plants and the <strong>crop</strong> management, including weed control actions<br />

(see also Table 4). The impacts of PFCs on arable weeds might therefore be caused by various<br />

mechanisms.<br />

1) PFCs are characterized by the absence of soil tillage during the whole duration of the <strong>crop</strong><br />

(about 2-6 years), thus often much longer than <strong>with</strong> annual <strong>crop</strong>s, where soil tillage and<br />

sowing operations are mostly effectuated once or even several times per year. This may<br />

have various impacts on weeds such as a reduced germination of weed species needing<br />

light or oxygen stimulus for germination (Huarte and Arnold, 2003) and an increased<br />

survivorship of established weed plants.<br />

2) In contrast to these reduced soil disturbances, mowing or grazing may lead to frequent<br />

mechanical disturbances of the aboveground vegetation. While most annual <strong>crop</strong>s are<br />

harvested only once per year, forage <strong>crop</strong> cutting is effectuated about 2-5 times per year,<br />

thus often both at earlier and later times of the year compared to the single harvesting date<br />

of annual <strong>crop</strong>s. This may reduce the survivorship, biomass and seed production of weeds<br />

(Gill and Holmes, 1997), although species may strongly differ in their sensitivity to<br />

cutting.<br />

3) PFCs are often characterized by strong canopy closures and <strong>de</strong>ep and <strong>de</strong>nse rooting<br />

systems. This may cause intense competition against weeds. Compared to annual <strong>crop</strong>s,<br />

competition may not only be stronger, but also more exten<strong>de</strong>d in time. While annual <strong>crop</strong>s<br />

are often characterized by periods of weak competition (i) during <strong>crop</strong> establishment, (ii)<br />

<strong>crop</strong> ripening/senescence and (iii) after harvest, perennial <strong>crop</strong>s have only one<br />

establishment phase in the first year and regrowth after cutting may be faster than initial<br />

27

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