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Diversifying crop rotations with temporary grasslands - Université de ...

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(illustrated in Fig. 24). Later, remobilisation will <strong>de</strong>crease again, which might be either due to<br />

a reduced <strong>de</strong>mand (sufficient photosynthesis on newly established leaves) and/or due to a<br />

reduced offer (<strong>de</strong>pletion of the belowground resources).<br />

Fig. 24: Daily aboveground biomass increase (ΔBMj)<br />

before and after a partial <strong>de</strong>struction of the<br />

photosynthetically active plant organs (cutting).<br />

ΔBMphsynth<br />

j, daily aboveground biomass increase powered by photosynthesis; ΔBMremob j,<br />

daily aboveground<br />

biomass increase powered by remobilisation of belowground carbohydrate resources. The arrow () indicates<br />

the amount of biomass produced by photosynthesis on the residual green surface remaining after cutting.<br />

ΔBMphsynth j <strong>de</strong>pends essentially on the plant’s green surface and may be simulated by the<br />

FLORSYS mo<strong>de</strong>l in the same way for uncut and cut plants (based on the energy balance of<br />

the plant estimated as a function of its light environment). In contrast, a new formula must be<br />

<strong>de</strong>veloped to estimate ΔBMremob j that takes into account the consi<strong>de</strong>rations and experimental<br />

ΔΒM<br />

ΔBMj<br />

remob j<br />

Species 2<br />

results discussed above:<br />

ΔBMphsynth j<br />

↓ Cutting<br />

LAth<br />

−LAj<br />

1 ⎛<br />

⎞<br />

( 0;<br />

T − T ) ⋅ ⋅⎜β<br />

⋅ BM − ∆BM<br />

⎟<br />

= ⋅ ⋅<br />

∑<br />

⎝<br />

j<br />

ΒΜbefore<br />

α max mean j b<br />

before<br />

LAth<br />

1<br />

The ‘remobilization of carbohydrates during day j’ (ΔBMremob j)<br />

would thus <strong>de</strong>pend on:<br />

• the aboveground biomass before cutting, BMbefore<br />

(that is correlated to belowground<br />

biomass for a given species, Gedroc et al., 1996; Wardle et al., 2004);<br />

<br />

174<br />

ΔBMremob j<br />

time<br />

ΔBMphsynth j<br />

remob j<br />

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