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Diversifying crop rotations with temporary grasslands - Université de ...

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<strong>crop</strong> treatments, weed emergence was even more reduced in the Dactylis <strong>crop</strong>s even if this<br />

<strong>crop</strong> showed higher initial weed <strong>de</strong>nsities and higher seed production. In this <strong>crop</strong>, the soil<br />

surface was generally har<strong>de</strong>r and often intertwined <strong>with</strong> fine and <strong>de</strong>nse grass roots, while the<br />

soil surface in the Medicago <strong>crop</strong>s was looser and often wetter, and comprised less superficial<br />

roots (Medicago roots are known to grow <strong>de</strong>eper than Dactylis grass roots), which might have<br />

favoured weed germinations.<br />

C.II.4.3.2 Competition (B)<br />

Competition for common growth resources (light, water, nutrients) and space is probably the<br />

most important direct interaction between <strong>crop</strong>s and weeds. An established and <strong>de</strong>nse plant<br />

canopy can affect several important stages of the weed life cycle (see Fig. 20 and Table 13 for<br />

an overview). Germination rates may be reduced due to a lack of light stimuli and a modified<br />

light quality un<strong>de</strong>r the canopy (Huarte and Arnold, 2003). Initial seedlings growth may be<br />

affected due to a reduced availability of water and nutrients or allelopathic interactions (Xuan<br />

and Tsuzuki, 2002). After seedling emergence, competition (for light) is probably the most<br />

important factor <strong>de</strong>termining weed growth, biomass and seed production in arable fields of<br />

temperate <strong>crop</strong>ping systems. There are several indications that competitive interactions played<br />

an important role for weed (and <strong>crop</strong>) growth in our experiment:<br />

• When pooling all <strong>crop</strong> treatments, <strong>crop</strong> and weed biomass showed strong negative<br />

correlations for all measurement dates during the first year of the experiment suggesting a<br />

strong impact of competition (Fig. 19). In particular, the initial weed biomass was high in<br />

all treatments and plots, where the initial <strong>crop</strong> biomass and competitive ability was low.<br />

This was the case in the cocksfoot <strong>crop</strong>s, which generally showed a slower initial<br />

establishment than alfalfa and winter wheat (Fig. 18). Moreover, the autumn-sown<br />

perennial <strong>crop</strong>s showed a better establishment than the spring-sown perennial <strong>crop</strong>s. When<br />

these two ‘unfavourable factors’ were combined (spring-sown cocksfoot <strong>crop</strong>s), initial<br />

weed biomass was highest (Fig. 18) and in turn reduced <strong>crop</strong> growth due to competition<br />

during the first month. However, the initial competitive disadvantages of spring sowing<br />

and of Dactylis <strong>crop</strong>s disappeared already during the first year ( Fig. 18).<br />

103

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