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Diversifying crop rotations with temporary grasslands - Université de ...

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Table 12: Factors differing between five <strong>crop</strong> treatment comparisons (as in).<br />

Factor<br />

Crop type Species<br />

96<br />

Treatment comparison<br />

Sowing<br />

season<br />

Cutting<br />

frequency<br />

Tillage in<br />

inter<strong>crop</strong><br />

Soil disturbance (superficial tillage, sowing)* An >> P<br />

D >~ M<br />

(roots?)<br />

Season = T+ > T-<br />

Aboveground vegetation disturbance<br />

(hay cuttings /<strong>crop</strong> harvesting)*<br />

P > An = A >~ Sp C+ >> C- T+ >~ T-<br />

Fertilisation (N-fertilizer vs. N-fixation) ≠ ≠ = = =<br />

Vegetation cover (competition for light)*<br />

≠ ≠ growth<br />

dynamics<br />

≠ growth<br />

dynamics<br />

≠ ≠ <strong>crop</strong><br />

establishm.<br />

C- > C+ =<br />

Allelopathy ?? ?? = = =<br />

* Impacts of these factors on the weed life cycle are <strong>de</strong>tailed in Table 13.<br />

C.II.4.1.1<br />

Plant <strong>de</strong>nsities<br />

Emerged plant <strong>de</strong>nsities and biomasses of many common annual weed species showed stable<br />

or <strong>de</strong>creasing ten<strong>de</strong>ncies in all perennial <strong>crop</strong> treatments. Such weed population dynamics<br />

may be surprising given the complete absence of soil tillage and specific weed control<br />

techniques in the PFCs. (Possible mechanisms of these <strong>de</strong>clines will be discussed below).<br />

Interestingly, reductions in weed abundances in PFCs were much more pronounced than<br />

reductions in species richness leading to improved diversity/<strong>de</strong>nsity-ratios ( Fig. 12) that may<br />

be useful for reducing the ‘weeds tra<strong>de</strong>-off’ (see § A.III.3 in the general introduction).<br />

In contrast, some weed species showed strongly increasing population sizes in the succession<br />

of annual <strong>crop</strong>s, as indicated by the increasing field emergence <strong>de</strong>nsities (Table 10, Fig. 12,<br />

Fig. 13) and weed biomasses (Fig. 17) during the 2.5 years experiment compared to the<br />

perennial <strong>crop</strong>s. These strong weed increases in the annual <strong>crop</strong>s are probably due to events of<br />

weed seed multiplication during the experimental period. It can not be explained by <strong>de</strong>nse<br />

background seed <strong>de</strong>nsities or the initial weed seed addition, which would be visible in all<br />

treatments of the randomized block experiment, which was not the case for any of the 16 sown<br />

species. In both spring-sown perennial <strong>crop</strong>s, consi<strong>de</strong>rable weed seed production occurred<br />

probably during the first month after <strong>crop</strong> sowing due to the slow initial <strong>crop</strong> establishment<br />

and the late first cutting (see discussion on the mechanism below).<br />

C.II.4.1.2<br />

Species composition<br />

Due to these heterogeneous weed population dynamics, species composition varied most<br />

strongly between annual and perennial <strong>crop</strong>s at the end of the experiment, ( Table 9). After the

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