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216<br />

Petals linear, 10–12 × ca. 1 mm, margins revolute, deflexed<br />

lower than lateral sepals but upcurved from middle; lip recurved<br />

abruptly at middle, cuneate-obovate, 10–12 × 7–10 mm,<br />

base contracted and without a callus, margin entire to indistinctly<br />

serrulate, apex truncate-rounded, minutely apiculate.<br />

Column curved, 5–6 mm, base dilated into 2 rounded mounds,<br />

apex with obtuse wings. Fl. Jun–Jul.<br />

● Forests; 1800–2000 m. E and N Taiwan.<br />

This species may be conspecific with Liparis makinoana Schlechter,<br />

from Japan, given that the descriptions of the two coincide. In<br />

Japan, the name L. makinoana has been misapplied to different taxa<br />

(e.g., by Tsutsumi & Yukawa, Bull. Natl. Mus. Nat. Sci., Tokyo, B, 34:<br />

89–94. 2008).<br />

6. Liparis campylostalix H. G. Reichenbach, Linnaea 41: 45.<br />

1877.<br />

羊耳蒜 yang er suan<br />

Leptorkis campylostalix (H. G. Reichenbach) Kuntze;<br />

Liparis giraldiana Kraenzlin; L. pauciflora Rolfe; L. yuana<br />

Ormerod.<br />

Herbs, terrestrial, rarely epiphytic. Pseudobulbs clustered,<br />

ovoid to globose, 5–12 × 3–8 mm, enclosed by 2 or 3 membranous<br />

sheaths. Leaves 2; petiole 1.5–8 cm, base sheathing, not<br />

articulate; blade ovate or ovate-oblong to subelliptic, (2–)5–<br />

10(–16) × (1–)2–4(–7) cm, base contracted into petiole, margin<br />

entire, occasionally wavy, apex acute to obtuse. Inflorescence<br />

10–<strong>25</strong> cm; peduncle 3.5–15 cm, winged; rachis 2–17 cm, laxly<br />

to subdensely few to many flowered; floral bracts lanceolate, 1–<br />

5.5 mm, apex acute. Flowers greenish, often tinted pink to purple<br />

or light purple; pedicel and ovary 5–10 mm. Dorsal sepal<br />

ligulate-lanceolate, 5–9 × 1.8–2 mm, 3-veined, apex subacute;<br />

lateral sepals parallel below lip, obliquely oblong-lanceolate,<br />

4.5–8.5 × 1.5–2 mm, 3-veined, apex subacute. Petals deflexed,<br />

parallel to and below lateral sepals, linear, 5–7 × ca. 0.5 mm; lip<br />

cuneate to oblong-obovate, 5–6 × 3–3.5 mm, base contracted,<br />

without a callus, margin entire to irregularly suberose, apex<br />

broadly apiculate, truncate to emarginate. Column weakly<br />

curved, 2.5–3.5 mm, base dilated, apex with small subquadrate<br />

wings. Fl. Aug–Oct.<br />

Forests, thickets, grasslands; 1100–2800(–3400) m. Gansu, Guizhou,<br />

Hebei, Heilongjiang, Henan, W Hubei, Jilin, Liaoning, Nei<br />

Mongol, Shandong, S Shanxi, Sichuan, Taiwan, Xizang, Yunnan [Japan,<br />

Korea, Russia (Far East)].<br />

Tsutsumi and Yukawa (Bull. Natl. Mus. Nat. Sci., Tokyo, B, 34:<br />

89–94. 2008) argue that the name Liparis japonica (Miquel) Maximowicz<br />

is actually a synonym of Malaxis monophyllos. The next available<br />

name for the “L. japonica” of authors is L. campylostalix. We have<br />

not followed Lang and Tsi (in Wu, Fl. Xizang. 5: 842–844. 1987) in<br />

separating “L. japonica” (larger plants, entire lip) from L. campylostalix<br />

(small plants, entire lip) given that intermediates exist.<br />

7. Liparis auriculata Blume ex Miquel, Ann. Mus. Bot.<br />

Lugduno-Batavi 2: 203. 1866.<br />

玉簪羊耳蒜 yu zan yang er suan<br />

Leptorkis auriculata (Blume ex Miquel) Kuntze; Liparis<br />

yakusimensis Masamune.<br />

Herbs, terrestrial, often forming large clusters. Pseudo-<br />

EPIDENDROIDEAE<br />

bulbs borne 3–5 cm apart, ovoid, slightly compressed, 1–2 × ca.<br />

1.5 cm. Leaves 2; petiole sheathlike, 4–6 cm, not articulate;<br />

blade ovate, broadly ovate, or cordate, 4–10 × 3–8 cm, base<br />

rounded to cordate and decurrent into petiole, apex acute. Inflorescence<br />

20–30 cm; rachis several to more than 10-flowered;<br />

floral bracts lanceolate, 1.5–2.5 mm. Flowers ca. 1 cm in diam.,<br />

greenish, purplish, or deep purplish red; pedicel and ovary 5–6<br />

mm. Dorsal sepal linear, 6–7 × 1.5–2 mm, apex acute; lateral<br />

sepals similar to dorsal sepal, slightly shorter and broader. Petals<br />

subfiliform, ca. 6 × 0.5 mm; lip orbicular or ovate-orbicular,<br />

5.5–6 × ca. 5 mm, with 2 subtriangular small calli near base,<br />

apex rounded or sometimes apiculate. Column recurved at apex,<br />

3–5 mm, slender, narrowly winged, base dilated. Fl. May–Jul.<br />

Dense forests, damp places; 1000–<strong>25</strong>00 m. C and N Taiwan<br />

[Japan (Ryukyu Islands)].<br />

8. Liparis cathcartii J. D. Hooker, Hooker’s Icon. Pl. 19: ad t.<br />

1808. 1889.<br />

二褶羊耳蒜 er zhe yang er suan<br />

Leptorkis cathcartii (J. D. Hooker) Kuntze.<br />

Herbs, terrestrial. Pseudobulbs ovoid, 5–6 × 4–5 mm, ±<br />

enclosed by white membranous sheaths. Leaves 2; petiole<br />

sheathlike, 2–5.5 cm, amplexicaul, not articulate; blade elliptic,<br />

ovate, or ovate-oblong, 3.5–8 × 1.7–4 cm, base contracted and<br />

decurrent into petiole, margin slightly crisped or nearly entire,<br />

apex acute or obtuse. Inflorescence 7–<strong>25</strong> cm; peduncle slightly<br />

compressed cylindric, narrowly winged on either side; rachis<br />

several to more than 10-flowered; floral bracts ca. 1 mm. Flowers<br />

pink, occasionally green or purple; pedicel and ovary 7–8<br />

mm. Sepals narrowly oblong, 7–9 × ca. 2.5 mm, inconspicuously<br />

3-veined, apex obtuse; lateral sepals slightly oblique. Petals<br />

nearly filiform, 7–9 × ca. 0.4 mm, 1-veined; lip obovate to<br />

elliptic-obovate, 8–9 × 7–8 mm, base contracted, usually with 2<br />

short longitudinal lamellae, very rarely lamellae inconspicuous,<br />

margin irregularly toothed, apex subtruncate and mucronate.<br />

Column arcuate, 3–3.5 mm, base dilated and thick, apex<br />

winged. Capsule obovoid-oblong, 1.1–1.3 cm × ca. 5 mm;<br />

fruiting pedicel 6–9 mm. Fl. Jun–Jul, fr. Oct.<br />

Humid or grassy places along valleys; 1900–<strong>25</strong>00 m. Sichuan, C<br />

and NW Yunnan [Bhutan, NE India, Nepal].<br />

9. Liparis pauliana Handel-Mazzetti, Anz. Akad. Wiss. Wien,<br />

Math.-Naturwiss. Kl. 58: 65. 1921.<br />

长唇羊耳蒜 chang chun yang er suan<br />

Liparis cucullata S. S. Chien.<br />

Herbs, terrestrial. Pseudobulbs ovoid or ovoid-oblong, 1–<br />

2.5 cm × 8–15 mm, ± enclosed by many white membranous<br />

sheaths. Leaves 2, very rarely 1; petiole sheathlike, 0.5–4 cm,<br />

amplexicaul, not articulate; blade ovate to elliptic, 2.7–9 × 1.5–<br />

5 cm, membranous or herbaceous, base contracted into petiole,<br />

margin crisped and irregularly serrate, apex acute or shortly<br />

acuminate. Inflorescence 7–28 cm; peduncle flattened cylindric,<br />

narrowly winged on either side; rachis laxly several flowered,<br />

very rarely many flowered or 1- or 2-flowered; floral<br />

bracts ovate or ovate-lanceolate, 1.5–3 mm. Flowers pale purple,<br />

sepals often pale yellowish green; pedicel and ovary 1–1.8

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