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FOC_25_Epidendroidea..

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458<br />

EPIDENDROIDEAE<br />

suborbicular; rostellum large, raised, laterally compressed, fleshy, with a longitudinal furrow along its edge into which stipe and<br />

dorsally placed pollinia recline; anther cap subglobose, apex beaked; pollinia 4, as 2 unequal pollen masses, waxy, compressed<br />

globose, separate, each with a short caudicle, attached to a common long stipe and a small, ovate viscidium.<br />

About 11 species: from China and India to Indonesia; two species (one endemic) in China.<br />

1a. Inflorescence longer than leaves; anther cap ca. 3 mm, longer than wide, apex narrowed and long beaked ............ 1. S. smithiana<br />

1b. Inflorescence shorter than leaves; anther cap ca. 1.5 mm, wider than long, apex not long beaked ....................... 2. S. magnirostris<br />

1. Sarcoglyphis smithiana (Kerr) Seidenfaden, Opera Bot. 114:<br />

383. 1992 [“smithianus”].<br />

大喙兰 da hui lan<br />

Sarcanthus smithianus Kerr, J. Siam Soc., Nat. Hist. Suppl.<br />

9: 239. 1933; Sarcoglyphis yunnanensis Z. H. Tsi.<br />

Stems 2–5 cm, enclosed in persistent leaf sheaths. Leaves<br />

narrowly oblong or slightly falcate-oblong, 11–19 × 1.5–2 cm,<br />

obtuse and unequally bilobed. Inflorescence racemose, rarely<br />

paniculate, pendulous, longer than leaves, many flowered; peduncle<br />

with 3 or 4 bractlike sheaths; rachis slender; floral bracts<br />

ovate, small, ca. 1 mm, acute. Flowers small, white, tinged with<br />

purple, lip purple; pedicel and ovary 5–8 mm. Dorsal sepal<br />

oblong, cymbiform, ca. 4.5 × 3 mm, obtuse; lateral sepals<br />

obliquely ovate, ca. 4.5 × 3.5 mm, obtuse. Petals oblong, ca. 4 ×<br />

1.2 mm, obtuse; lip spurred, 3-lobed; lateral lobes erect, subtriangular,<br />

apex rounded; mid-lobe slightly fleshy, middle dilated<br />

and transversely oblong, beaked toward apex; spur subconic,<br />

ca. 3 mm, apex obtuse, interior slightly septate, with a laterally<br />

compressed callus on back wall; callus deeply grooved centrally,<br />

with 2 small hornlike projections, base densely minutely<br />

papillate. Column ca. 1.5 mm; rostellum ± oblong, ca. 1.5 mm.<br />

Fl. Apr.<br />

Epiphytic on tree trunks in evergreen broad-leaved forests; 500–<br />

700 m. S Yunnan [Laos, Thailand, N Vietnam].<br />

2. Sarcoglyphis magnirostris Z. H. Tsi, Acta Phytotax. Sin. 23:<br />

387. 1985 [“Sarcoglyphia”].<br />

短帽大喙兰 duan mao da hui lan<br />

163. CLEISOSTOMA Blume, Bijdr. 362. 18<strong>25</strong>.<br />

隔距兰属 ge ju lan shu<br />

Chen Xinqi (陈心启 Chen Sing-chi); Jeffrey J. Wood<br />

Stems ascending, 1–4 cm, ca. 1 cm in diam., usually 3–5leaved.<br />

Leaves oblong, usually 14–16 × 1.8–2 cm, unequally bilobed.<br />

Inflorescence pendulous, racemose, shorter than leaves,<br />

unbranched, sparsely many flowered; rachis purplish black;<br />

floral bracts ovate-triangular, ca. 1 mm, acute. Flowers whitish,<br />

tinged with purplish red or purple, spur white, rostellum yellowish<br />

white, apex of anther cap yellowish brown with 2 purple<br />

stripes; pedicel and ovary pale purple, ca. 1 cm. Dorsal<br />

sepal ovate-elliptic, 4–5 × 3–3.5 mm, rounded; lateral sepals<br />

obliquely ovate, slightly larger than dorsal sepal, obtuse. Petals<br />

oblong, ca. 4 × 2 mm, obtuse; lip slightly fleshy, 3-lobed; lateral<br />

lobes erect, nearly suborbicular; mid-lobe broadly ligulate, ca. 3<br />

× 3.2 mm, concave at center above, apex subtruncate; spur subconic,<br />

ca. 3 mm, apex rounded, inside slightly septate, with a<br />

callus on back wall; callus subquadrate, grooved centrally. Column<br />

stout; rostellum suborbicular, much higher than clinandrium;<br />

anther cap nearly triangular, ca. 1.5 mm. Fl. May.<br />

● Epiphytic on tree trunks in forests along valleys; ca. 800 m. S<br />

Yunnan.<br />

Sarcanthus Lindley (1826), not Lindley (1824).<br />

Herbs, epiphytic or rarely lithophytic, monopodial, small to medium-sized. Stems erect or pendulous, rarely creeping, long or<br />

short, rigid, branched or unbranched, many noded. Leaves few to many, distichous, dorsiventrally flattened or terete, base usually<br />

jointed and sheathing, apex bilobed or acute. Inflorescence racemose or paniculate, lateral, axillary or opposite leaves, erect, horizontal,<br />

or pendulous, few to many flowered; floral bracts small, much shorter than pedicel and ovary. Flowers small, ± fleshy. Sepals<br />

free, usually spreading. Petals free, often smaller than sepals; lip adnate to column by its base or back margins of lateral lobes, immovable,<br />

spurred at base, 3-lobed; disk often with longitudinal lamellae or ridges; spur conic, cylindric, or saccate, often longitudinally<br />

septate inside, rarely septum reduced to a narrow keel, usually with a callus on back wall and often with outgrowths on front<br />

wall closing entrance. Column short, stout, foot short; rostellum small; pollinia 4, appearing as 2 unequal masses, waxy, subglobose,<br />

with a common stipe and a solitary small and subglobose to broad and horseshoe-shaped viscidium.<br />

About 100 species: Sri Lanka, India, mainland SE Asia, Malaysia, and Indonesia to the Philippines, New Guinea, Pacific islands, and Australia;<br />

16 species (four endemic) in China.<br />

1a. Leaves cylindric or subcylindric, terete, 2–3 mm in diam.<br />

2a. Leaves 6–11 cm.<br />

3a. Column with apical stelidia, glabrous; stipe broadly ovate-triangular or miter-shaped; viscidium<br />

crescent-shaped ......................................................................................................................................... 12. C. williamsonii<br />

3b. Column without stelidia, densely hairy at base in front; stipe nearly suborbicular; viscidium<br />

U-shaped .......................................................................................................................................................... 15. C. simondii<br />

2b. Leaves 17–33 cm.

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