Bulletin of the British Museum (Natural History)

LICHEN GENUS MICAREA IN EUROPE 199
placed in Trapelia Choisy or Trapeliopsis Hertel & G. Schneider, or if a new genus is required to
accommodate them.
Lecidea littorella Nyl. in Flora, Jena 60: 229 (1877). - Catillaria littorella (Nyl.) Zahlbr., Cat. lich. univ. 4:
56 (1926). Type: Ireland, West Galway, 'Hibernia occidentalis, Bord du Lough Inagh, mais extremement
rare', 1876, C. Larbalestier (Yi-^Yl. 18884- holotype!, isotypes: BM!, BM ex K!).
The following notes were made from the holotype: Thallus saxicolous, not delimited but forming small
patches amongst Hymenelia lacustris and Porina chlorotica, whitish, or dull ochraceous in part (? due to
age), rimose, matt. Apothecia numerous, weakly marginate, plane to slightly convex, pallid to dull
orange-red, 0-1-0-3 mm diam; margin not exceeding the level of the disc, very thin, c. 0-1-0-3 mm diam;
margin not exceeding the level of the disc, very thin, c. 0-01-0-02 mm wide, whitish with faintly pruinose
appearance. Hymenium 40-47 jxxn tall, hyaline. Asci cylindrical-clavate, ' Lecanora-type\ 8-spored.
Spores ovoid to ovoid-oblong, often 'slipper-shaped', 1-septate and often constricted at the septum,
9-5-14(-16)x4-6(-7) ^tm (Fig. 56B). Paraphyses numerous, thin, 0-8-1 ixm wide, simple or sparingly
branched below, becoming wider (to 1-7 ixm) and more frequently branched above. Hypothecium hyaline.
Excipulum hyaline or dilute yellow-straw, inspersed with minute crystals in water mounts but ± clearing in
K, c. 20 /am wide laterally, slightly widening below to 28 )u,m, minutely paraplechtenchymatous with cells in
the lower part measuring 4—6x2-5^ /u,m. Pycnidia frequent, at first immersed, later becoming emergent,
white (translucent when wet), c. 80-100 fxm diam. Conidia (Fig. 56C) simple, ovoid to pyriform,
2-8xl-5-l-7)Ltm.
L. littorella was provisionally referred to Micarea by Kilias (1981: 392). However, my subsequent
examination of the holotype proved it to be a saxicolous form of the normally foliicolous Catillaria
bouteillei (Desm.) Zahlbr.; previous reports of this species on rock are given by Degelius (1944) and
Santesson (1952). C. bouteillei is apparently widely distributed in Ireland, especially on the leaves oiBuxus
(Knowles, 1929; Scannell, 1978). An additional British saxicolous specimen has been located: V.C.5,
South Somerset, Broomfield, Ruborough Camp, 1914, W. Watson (BM ex K).
Nylander's choice of the epithet 'littorella' is rather misleading because the holotype was not collected on
the sea-shore, but on the shores of an inland, freshwater lake some 10 km from the sea. British authors (e.g.
James, 1970; Fletcher, 1975) have mistakenly applied the name Catillaria littorella to a species that grows in
crevices in siliceous rocks near the sea-shore; this species probably belongs to the perplexing Lecania
ery5/fte complex.
Lecidea milliaria a. [var.] terrestrisFr. Lich. Eur. 342 (1831). Type: not designated.
This name is of unlikely appHcation in M/carea. Fries cited his L/c/i. Suec. no. 213, the example of which
in UPS is a member of the Lecidea limosa group.
Lecidea ocelliformis Nyl. in Flora, Jena 48: 145 (1865). - Bilimbia ocelliformis (Nyl.) Branth. & Rostr. in
Bot. Tidskr. 3: 226 (1869). - Lecidea atroviridis f. ocelliformis (Nyl.) Hedl. in Bih. K. svenska
VetenskAkad Handl. Ill, 18 (3): 64 (1892). - Catillaria prasina f. ocelliformis (Nyl.) Erichsen in Annls.
mycol. 41: 205 (1943). Type: Finland, Tavastia australis, Hollola, ad corticem Sorbi, 30 vi 1863, /. P.
Norrlin 210 (H-NYL 20607! - labelled Tsotype' by M. Inoue in 1979).
This is not a Micarea. If the types of L. atroviridis (Arnold) Th. Fr. and L. ocelliformis are conspecific, as
placed by Hedlund (op. cit.) and Vainio (1934: 218), then the latter epithet has priority.
Lecidea pauxilla Krempelh. in Verh. zool.-bot. Ges. Wien 26: 455 (1876). Type: New Zealand, on bark
[? Podocarpus], 18 -, C. Knight (M - lectotype!; M - isolectotype!).
L. pauxilla was given as a synonym of Catillaria synothea (i.e. Micarea denigrata) by Zahlbruckner (Cat.
lich. univ. 4: 78, 1926). However, the type material belongs to Cliostomum griffithii (Sm.) Coppins.
Lecidea recondita Erichsen in Annls mycol. 42: 25 (1944). Type: Germany, Schleswig-Holstein: 'Apen-
rade, an Blocken am Grunde einer Erosionsschluct im Geholz Jiirgensgaard' [now Denmark, Jylland,
near Abenra], 28 vii 1932, C. F. E. Erichsen (HBG - holotype!).
In the protologue Erichsen compares this with Lecidea sylvicola (= Micarea sylvicold). My examinations
of the holotype proved it not to be a Micarea but referable to Catillaria chalybeia (Borrer) Massal.; see
Kilias (1980: 448).
Lecidea sabuletorum f. simp/icior Nyl., Lich. Scand.: 205 (1861). - Bilimbia trisepta f. simplicior (Nyl.)
Vainio in Acta Soc. Fauna Fl. fenn. 53 (1): 258 (1922). Type: not designated.
Nylander (loc. cit.) cited the following material 'supra muscos ad Helsingforse [ipse] et in Sueciae
montibus [Thedenius] atque supra terram in Lapponia'.
According to Fries (1874: 523) the 'Helsingforse' specimen is Lecidea verrucula (= Micarea turfosa);
according to Vainio (1922: 140) the Swedish specimen from Funnesdalsberget in Harjedalen collected by