Bulletin of the British Museum (Natural History)

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34. Micarea prasina Fr. LICHEN GENUS MICAREA IN EUROPE 173 -^^r Map 17 Micarea peliocarpa # 1950 onwards O Before 1950 (Figs ID, 26-27, 49, 53-54; Maps 18-20) Syst. orb. : 256-7 (Dec. 1825). - Biatora prasina Fr. , Stirp. agrifemsion.: 36 (June 1825), nom. illeg. (Art. 63). - Biatora prasina (Fr.) Trevisan, Linnaea 28: 288 (1856), nee Tuck. & Mont., in Mont. (1857). - Catillaria prasina (Fr.) Th. Fr., Lich. Scand. 2: 572 (1874). See note (i) below and note on Sporacestra under 'Excluded taxa'. Biatora micrococca Korber, Parerg. lich.: 155 (1860). - Catillaria micrococca (Korber) Th. Fr., Lich. Scand. 2: 571 (1874). - Micarea prasina f. micrococca (Korber) Hedl. in Bih. K. svenska VetenskAkad. Handl. Ill, 18 (3): 77, 87 (1892). - Micarea micrococca (Korber) H. Gams, Kleine Kryptfl. 3: 67 (1967); comb, inval. (Art. 33.2). Type: Germany, Baden-Wiirttemberg, 'in regno Wurtembergico', on Pinus bark, 1862, K. A. Kemmler, Korber Lich. Sel. germ. 250 (L 910, 139-1361-neotype! [t.l.c: 'prasina- unknown A']). See note (ii) below. Lecidea subviridescens Nyl. in Flora, Jena 51: 474 (1868) . - Micarea subviridescens (Nyl.) Hedl. in Bih. K. svenska VetenskAkad. Handl. Ill, 18 (3): 77, 87 (1892). - Bacidia subviridescens (Nyl.) Zahlbr., Cat. lich. univ. 4: 154 (1926). Type: Channel Islands, Jersey, Boulay Bay, on coastal turf and soil, 1868, C. Larbalestier (H-NYL 19056 [as 'subvirescens'] - lectotype! ; BM - isolectotype! [t.l.c. : 'prasina unknown C']; BM-topotypes!). See note (iii) below. Lecidea prasiniza Nyl. in Flora, Jena 57: 312 (1874). - Micarea prasina f. byssacea subf . prasiniza (Nyl.) Th. Fr. in Hedl. in Bih. K. svenska VetenskAkad. Handl. Ill, 18 (3): 77 (1892). Type: Finland, Tavastia australis, Padasjoki, Nyystola, on Alnus, 1872, E. A. Lang fVainio] (H-NYL 21604 - lectotype!, H - isolectotype!). [material insufficient for t.l.c.]. Lecidea sordidescens Nyl. in Flora, Jena 57: 312 (1874). - Lecidea erysiboides f. sordidescens Nyl. in Norrlin in Not. Sdllsk. Fauna Fl. fenn. Forh. 11: 188 (1870); nom nudum (Art. 32). - Catillaria prasina
174 BRIAN JOHN COPPINS var. byssacea f . sordidescens (Nyl.) Blomb. & Forss. , Enum. PI. Scand. : 91 (1880). - Micarea prasina f byssacea subf. sordidescens (Nyl.) Th. Fr. in Hedl. in Bih. K. svenska VetenskAkad. Handl. Ill, 18 (3): 77 (1892). - Lecidea byssacea var. sordidescens (Nyl.) Vainio in Termeszetr. Fuz. 22: 320 (1899). - Catillaria prasina var. sordidescens (Nyl.) Lettau in Hedwigia 52: 136 (1912). - Micarea prasina var. sordidescens (Nyl.) Brodo in Bull. N. Y. St. Mus. Sci. Serv. 410: 152 (1968). Type: Switzerland, Zurich, 'An faulen Fichten-Strunken Z. H.', Hepp, Flecht. Eur. 11^ (E - lectotype! [t.l.c.: 'prasina-unknown B']; isolectotypes: BM (boxed set)! [t.l.c: 'prasina-unknown B'], H-NYL 21632 [fragment]!, M!). See note (iv) below. Lecidea prasiniza var. prasinoleuca Nyl. in Flora, Jena 64: 7 (1881). Type: Germany, Baden-Wiirttemberg, Heidelberg, Konigstuhle, on Picea abies, ix 1880, Zwackh, Lich. Exs. 593a (H-NYL 21601 - lectotype! [t.l.c: 'prasina-unknown A']). See note (v) below. Catillaria micrococca f. glebosula Erichsen in Annls mycol. 36: 139 (1938). Type: Germany, Schleswig- Holstein, Eiderstedt, Ording, at base of Betula, 4 viii 1913, C. F. E. Erichsen (HBG - holotype! [t.l.c: 'prasina-unknown B']. Catillaria micrococca var. discrepans Erichsen in Annls mycol. 36: 139 (1938). Type: Denmark, Jylland, Abenra (Apenrade), on smooth bark oiAlnus in a valley near Elisenlund, 3 ix 1913, C. F. E. Erichsen (HBG - holotype!). [material insufficient for t.l.c]. Catillaria prasina var. occulta Erichsen in Annls mycol. 36: 140 (1938). Type: Germany, Schleswig- Holstein, Ratzeburg, in the wood 'Bak', on ± smooth bark oiAlnus, 21 iii 1918, C. F. E. Erichsen (HBG - holotype!). [material insufficient for t.l.c]. Lecidea declivitatum Erichsen in Mitt. Inst. allg. Bot. Hamb. 10: 408 (1939). Type: Germany, Schleswig- Holstein, 'Krs Eutin: an der Boschung eines Waldwegs an der Nordwest-Seite des Ugleisees', 5 vi 1914, C. F. E. Erichsen (HBG -holotype! [t.l.c: 'prasina-unknown C']). Micarea polytrichi Poelt & Dobb. in Bot. Jb. 96: 343 (1975). Type: Austria, Steiermark, Grazer Bergland, a little north of Maria Trost near the Wenisbucher Strasse, c. 460 m, on Polytrichumformosum on shady side of a narrow gorge, 30 iii 1974, /. Poelt (GZU - holotype!) [material insufficient for t.l.c]. Catillaria prasina a laeta Th. Fr., Lich. Scand. 2: 573 (1874); nom. inval. (Art. 26). ?Biatora byssacea Zwackh in Flora, Jena 45: 510 (1862), non Hampe in Linnaea 25: 709 (1852). Type: Germany, Heidelberg, 'Konigsstuhle', on bark of old Quercus, ? Zwackh (not traced). See note (vi) below. ?Catillaria melanobola f. biseptata B. de Lesd., Rech. Lich. Dunkerque: 198 (1910). Type: France, Nord, Dunkerque, St Pol, on a piece of leather in dunes, B. de Lesdain (not seen). ?Catillaria melanobola f. nigra B. de Lesd., Rech. Lich. Dunkerque: 199 (1910). Type: France, Nord, Dunkerque, lignum of hollow Salix, B. de Lesdain (not seen). Notes: (i) Biatora prasina Fr. (June 1825) is superfluous because the protologue includes the sentence: 'Crusta est Byss. botryoides Linn!'. Byssus botryoides L. is a valid name {Sp. pi. 2: 1169 (1753)) but the material in the Linnaean herbarium (LINN) is a specimen (1278.16) labelled by Ehrhart; it is an alga and bears a determination label: 'Protococcus viridis Ag. Determined by Francis Drouet iv. 1950'. In the protologue of Micarea and M. prasina. Fries (Dec. 1825) makes a less dogmatic statement, merely indicating that when sterile M. prasina is indistinguishable from B. botryoides; M. prasina is, thereby, not rendered superfluous. (ii) I have seen two specimens named by Korber and collected from 'Wiirtemberg' by Kemmler: an undated specimen in WRSL and possibly part of the original gathering, and a later specimen issued in Korber's exsiccata. The first specimen would have to be completely destroyed for t.l.c. analysis, whereas the exsiccate specimen is larger and was found to contain 'prasina-unknown A', and it is this specimen that I have chosen for typification. If future studies require this chemical race to be recognized at specific rank, then 'micrococca' is the earliest available epithet. (iii) L. subviridescens is the earliest name for the chemical race with 'prasina-unknown C. (iv) Although Nylander (1874) did not provide a description, he validated L. sordidescens by referring to it Hepp Flecht. Eur. 278; the printed label for this exsiccatum includes a short description and a drawing of spores. (v) If the chemical race containing 'prasina-unknown A' were to be awarded varietal rank, then this name provides the earliest available epithet, provided that 'byssacea' is not available; see also note (vi). (vi) I have not seen a holotype specimen or any material suitable for the lectotypification of Biatora byssacea Zwackh. A neotype has not been selected because this name is a later homonym of B. byssacea Hampe. Hampe's name was given to a specimen collected by Charles Stuart in Tasmania, and the protologue ends with 'an status degenerans'. Biatora byssacea Hampe is included by Zahlbruckner (Cat. lich. univ. 3: 897) in a list of excluded and dubious names and is not accorded a catalogue number; the type .
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LICHEN GENUS MICAREA IN EUROPE 95 T
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Infrageneric considerations LICHEN
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