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Bulletin of the British Museum (Natural History)

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LICHEN GENUS MICAREA IN EUROPE 145<br />

Map 9 Micarea lignaria var. lignaria # 1950 onwards O Before 1950<br />

plant debris, loose stones, pieces <strong>of</strong> timber and sack-cloth. Mr V. Giavarini has recently found it<br />

in Dorset, growing on waste heaps <strong>of</strong> slag clay, in <strong>the</strong> company <strong>of</strong> M. leprosula, Baeomyces<br />

roseus, Cladonia arbuscula, C. ciliata, C. furcata, C. portentosa, and Coeldcaulon aculeatum s.<br />

lat. M. lignaria is very rare in <strong>the</strong> lowlands <strong>of</strong> south-east England, but has been found <strong>the</strong>re<br />

growing directly on <strong>the</strong> rock <strong>of</strong> east or north facing sandstone walls and churchyard memorials,<br />

and also on natural sandstone outcrops <strong>of</strong> <strong>the</strong> Sussex Weald.<br />

In upland areas M. lignaria is frequently found on <strong>the</strong> exposed lignum <strong>of</strong> fallen trunks<br />

(especially <strong>of</strong> conifers) and old timberwork, with, for example, Cladonia spp., Hypogymnia<br />

physodes, Micarea peliocarpa, Mycoblastus sterilis, Ochrolechia turneri, Parmelia saxatilis, and<br />

Lecanora polytropa (worked timber) . Occurrences <strong>of</strong> M. lignaria on <strong>the</strong> bark <strong>of</strong> healthy trees<br />

are ra<strong>the</strong>r rare, and in Britain are confined to <strong>the</strong> high rainfall areas <strong>of</strong> <strong>the</strong> north and west, where<br />

it has been collected on Alnus, Betula, Fraxinus, Ilex, Quercus, and old Sambucus.<br />

Reports <strong>of</strong> M. lignaria on mosses on limestone rocks usually result from <strong>the</strong> misidentification<br />

<strong>of</strong> Bacidia sabuletorum or Toninia lobulata. However, on a few occasions I have seen M. lignaria<br />

growing on thick bryophyte cushions or mats at high altitudes over limestone in <strong>the</strong> north<br />

Pennines, and over calcareous mica-schist in <strong>the</strong> Breadalbane Mountains; in such situations <strong>the</strong><br />

pH and calcium content <strong>of</strong> <strong>the</strong> substratum is presumably kept low by <strong>the</strong> leaching effect <strong>of</strong> <strong>the</strong>'<br />

high rainfall in those areas.<br />

The altitude range <strong>of</strong> M. lignaria in Britain is from sea-level to about 1200 m (Ben Lawers),<br />

although it may well occur at higher altitudes in <strong>the</strong> Ben Nevis group and <strong>the</strong> Cairngorni<br />

Mountains. Higher altitudes are attained in <strong>the</strong> mountains <strong>of</strong> central Europe, from where it has

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