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Physico-Chemical and Molecular Markers for Resistance to Insect Pests 157<br />

TABLE 6.1<br />

Morphological Markers Associated with Resistance to Heliothis/Helicoverpa<br />

Crop Morphological Trait Remarks References<br />

Cotton Nectarless Nonpreference Lukefahr et al. (1960)<br />

Leaf pubescence Oviposition<br />

preference<br />

Butter et al. (1996)<br />

Low bract teeth Resistance Rao et al. (1996)<br />

Red leaf Resistance Ansingkar et al. (1984)<br />

Okra leaf Resistance Wu, Cai, and Zhang (1997)<br />

Frego-bract Resistance Wu, Cai, and Zhang (1997)<br />

Erect growth habit Resistance Navasero and Ramaswamy (1991)<br />

Glandless Susceptibility Narayanan and Singh (1994)<br />

Tough bolls Resistance Mohan et al. (1995)<br />

Pigeonpea Nonglandular trichomes Resistance Romeis, Shanower, and Peter (1999)<br />

Colored seeds Resistance Nanda, Sasmal, and Mohanty (1996)<br />

Long and wide pods Susceptibility Nanda, Sasmal, and Mohanty (1996)<br />

Clustered podding Susceptibility Raut et al. (1993)<br />

Indeterminate growth habit Resistance Lateef and Pimbert (1990)<br />

Chickpea Glandular trichomes Resistance Yoshida, Cowgill, and Wightman<br />

(1997)<br />

Long and wide pods Susceptibility Gururaj et al. (1993)<br />

Tomato Glandular trichomes Resistance Farrar and Kennedy (1988),<br />

Sivaprakasam (1996)<br />

Pubescence Oviposition<br />

preference<br />

AVRDC (1987)<br />

Calyx texture and fl owering Resistance Cosenza and Green (1979)<br />

Fruit skin Resistance Cosenza and Green (1979)<br />

Bigger vine size Resistance Ferry and Cuthbert (1973)<br />

Soybean Trichomes Resistance Panda (1979)<br />

Visual Stimuli<br />

Insects use vision to recognize the shape of, and chemical stimuli to perceive, the host<br />

plant. Host fi nding from a distance is the fi rst step in host plant selection and colonization<br />

by insects. It involves the visual stimuli emitted by the host plant, for example, aphids are<br />

attracted to yellow-white surfaces (Moericke, 1969; Prokopy and Owens, 1983), cotton white<br />

fl y, Bemisia tabaci (Genn.) is attracted to yellow-green and yellow-red (Hussain and Trehan,<br />

1940), and sorghum midge, Stenodiplosis sorghicola (Coquillett) to white-yellow colors<br />

(Sharma, Leuschner, and Vidyasagar, 1990; Sharma and Franzmann, 2002). Grasshoppers<br />

orient themselves towards vertical lines or objects (Williams, 1954). The glossy trait of<br />

cabbage, Brassica oleracea var capitata L. leaves confers antixenosis to the cabbage aphid,<br />

Brevicoryne brassicae L. (Singh and Ellis, 1993). Sorghum genotypes with glossy leaf trait<br />

(light, yellow, shiny, and erect leaves) (Figure 6.1) are nonpreferred for oviposition by sorghum<br />

shoot fl y, Atherigona soccata (Rondani) as compared to genotypes with dark green,<br />

drooping, and nonshiny leaves (Sharma and Nwanze, 1997). Plants having yellowish green<br />

leaves are nonpreferred for oviposition by pod borer, Helicoverpa armigera (Hubner) as

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