Checklist of the Terrestrial and freshwater invertebrate fauna ... - UNIS

The terrestrial and freshwater invertebrate fauna of 

Svalbard: a checklist 

06 June 2013 Page 1 of 185

Notes: This is not the definitative inventory! 

This list is based on a collection of over 600 articles concerning the terrestrial invertebrate fauna of Svalbard. Nonetheless it should be not be considered as an 

accurate overview of the complete invertebrate fauna of the archipelago. The terrestrial invertebrate fauna is largely only known from two localities on 

Svalbard, Ny-Ålesund (Kongsfjorden) and the Isfjorden region. Little is known about the fauna of the east coast or the interior. In addition there are large 

numbers of doubtful identifications and possible confusion in the taxonomy. 

Wherever possible synonyms occuring in the Svalbard literature are presented in smaller font. 

This list is an ongoing project of Coulson, S.J. 2007 The terrestrial and freshwater invertebrate fauna of the High Arctic archipelago of Svalbard. Zootaxa 

1448; 41-58 and Coulson, S.J. Refseth, D. (2004) The terrestrial and freshwater invertebrate fauna of Svalbard (and Jan Mayen), pp57-122 in A catalogue of 

the terrestrial and marine animals of Svalbard (eds. Prestrud, P. Strøm H. Goldman H.). Skrifter 201. Norwegian Polar Institute, Tromsø. 

Steve Coulson 

06 June 2013 


The Terrestrial and Freshwater Invertebrate Fauna of Svalbard: 

Sarcomastigophora 

Heliozoea 

_ 

_ 

_ 

Actinophryridae 

Actinophrys sol Ehrenberg 

Clathrulinidae 

Clathrulina elegans Cienk 

Zoomastigophorea 

Cercomonadida 

_ 

_ 

Cercomonadidae 

Cercomonas gigantica Myl'nikov 460 

Cercomonas kiaerdammane Myl'nikov 460 


Rhizopoda 

Filosea 

Gromiida 

_ 

_ 


Cyphoderiidae 

Cyphoderia ampulla Leidy 22 

Cyphoderia perlucidus Beyens and Charedez 21 22 

Euglyphidae 

Assulina muscorum Greeff 14 22 24 25 244 260 

Assulina muscorum var. denticulata Chardez 22 

Assulina muscorum var. stenostoma Schönborn 22 260 

Assulina seminulum (Ehrenberg) Leidy 14 260 319 



Euglypha acanthophora Perty 22 25 

Euglypha alveolata Dujardin 319 

Euglypha ciliata Ehrenberg 14 22 319 

Euglypha ciliata var. glabra Wailes 260 

Euglypha cristata Leidy 319 

Euglypha compressa Carter 22 260 

Euglypha filifera Penard 22 

Euglypha laevis (Ehrenberg) Perty 14 22 244 260 

Euglypha laevis var. lanceolata Playfair 22 



Euglypha minor Chardez 22 

Euglypha polylepsis Bonnet and Thomas 22 

Euglypha rotunda Wailes and Penard 14 21 22 24 25 260 

Euglypha rotunda var. dorsalis Decloitre 14 

Euglypha rotunda var. glabra Wailes 14 

Euglypha rotunda var. minor Wailes 14 

Euglypha strigosa Leidy 22 24 25 

Euglypha strigosa f. glabra Wailes 14 22 24 25 244 260 

Euglypha strigosa var. muscorum Wailes 22 24 



Euglypha tuberculata Dujardin 14 22 25 420 421 

Euglypha tuberculata var. subcylindrica (Playfair) Decloitre 14 

Tracheleuglypha dentata (Vejdovsky) Deflandre 244 

Valkanovia delicatula Valkanov 14 

Gromiidae 

Gromia sp. 319 

Nadinella tenella Penard 244 

Pseudodifflugia globulosa Stepanek 244 

Pseudodifflugia gracilis Schlumberger 22 244 

Pseudodifflugia gracilis var. terricola Bonnet and Thomas 244 



Pseudodifflugia senartensis Couteaux 244 

Paulinellidae 

Paulinella chromatophora Lauterborn 22 

Trinematidae 

Corythion aerophila Decoitre 24 

Corythion delamarei Bonnet and Thomas 14 

Corythion dubium Taranek 14 22 24 260 

Corythion dubium var. aerophila Decloitre 14 24 

Corythion dubium f. minima Chardez 22 24 

Corythion dubium var. orbicularis Penard 22 24 

Corythion dubium var. terricola Schönborn 244 



Corythion pulchellum Penard 14 

Trinema complanatum Penard 22 25 

Trinema complanatum var. globulosa Chardez 22 

Trinema enchelys (Ehrenberg) Leidy 14 22 24 25 260 319 

Trinema lineare Penard 14 21 22 25 244 260 

Trinema lineare f. minuscula Chardez 14 22 

Trinema lineare f. truncatum Chardez 22 

Trinema lineare var. longa Korganova 22 

Trinema navicularis Decloitre 22 



Trinema penardi Thomas and Chardez 22 


Lobosea 

Amoebida 

_ 

_ 


Amoebidae 

Amoeba radiosa Ehrenberg 

Amoeba verrucosa Ehrenberg 

Arcellinida 

_ 

_ 

Arcellidae 

Arcella atrocera Leidy 319 

Arcella arenaria Greeff 22 25 260 

Arcella arenaria f. compressa Chardez 22 24 

Arcella arenaria var. sphagnicola Deflandre 22 



Arcella bathystoma Deflandre 22 

Arcella discoides Ehrenberg 22 244 

Arcella ovaliformis Chardez and Beyens 22 61 

Arcella rotundata Playfair 22 24 25 

Arcella rotundata var. aplanata Deflandre 22 

Arcella rotundata var. undulata Stepanek 22 25 

Arcella vulgaris Ehrenberg 14 420 421 

Chlamydophrys stercorea Cienk 319 

Cochliopodium erinaceum Penard 14 



Cochliopodium obscurum Penard 22 25 

Cochliopodium opalinum Penard 22 25 

Difflugiella horrida Schönborn 14 

Difflugiella oviformis (Penard) Bonnet and Thomas 14 

Difflugiella pusilla (Playfair) Grospietsch 

Microcorycia flava Greef 22 25 

Microcorycia husvikensis Beyens and Chardez 23 

Parmulina louisi Chardez and Beyens 22 

Pseudochlamys patella C. and L. 319 



Pyxidicua cymbalum Penard 25 

Pyxidicula ornata Bartos 14 

Centropyxidae 

Centropyxis aculeata Stein 22 25 244 

Centropyxis aculeata f. compressa 244 

Centropyxis acuminata Couteaux and Chardez 22 

Centropyxis aerophila Deflandre 14 21 22 24 25 244 260 

Centropyxis aerophila var. constricta Decloitre 22 

Centropyxis aerophila var. deflandrei Decloitre 22 

Centropyxis aerophila var. sphagnicola Deflandre 14 22 24 25 244 260 



Centropyxis aerophila var. minuta Chardez 24 25 

Centropyxis cassis (Wallach) Deflandre 14 22 

Centropyxis cassis var minima van Oye 22 420 421 

Centropyxis constricta (Ehrenburg) Penard 14 

Centropyxis cordiformis Couteaux 22 

Centropyxis delicatula Penard 22 

Centropyxis delicatul var. maior Jung 22 

Centropyxis ecornis Leidy 22 

Centropyxis ecornis var. deflandrei Thomas 14 



Centropyxis gasparella 516 

Centropyxis gasparella var. corniculata Beyens and Chardez 516 

Centropyxis gibba var. gibbosa Godeanu 21 22 23 

Centropyxis globulosa Bonnet and Thomas 22 

Centropyxis laevigata var. maior Jung 22 

Centropyxis lata Jung 22 

Centropyxis minima Deflandre 22 

Centropyxis minuta Deflandre 22 

Centropyxis orbicularis Deflandre 14 22 



Centropyxis plagiostoma Bonnet and Thomas 21 22 260 

Centropyxis platystoma (Penard) Deflandre 14 22 244 260 420 421 

Centropyxis pontigulasiformis Beyens and Chardez 21 22 244 

Centropyxis sylvatica Thomas 21 22 24 244 

Centropyxis sylvatica var. minor Bonnet and Thomas 22 

Centropyxis ventricosa Bartos 22 

Cryptodifflugiidae 

Cryptodifflugia compressa f. angustioris Tarnogradsky 22 24 

Difflugiidae 

Difflugia bacillifera Penard 22 

Difflugia bryophila Jung 21 22 24 244 



Difflugia compressa var. minima Schönborn 244 

Difflugia constricta Ehrenberg 319 

Difflugia dujardini Chardez 21 22 

Difflugia elegans Penard 244 

Difflugia elegans var. parva Chardez 244 

Difflugia glans Penard 22 24 

Difflugia globularis Leidy 21 22 

Difflugia globulosa Dujardin 22 24 260 319 

Difflugia globulus Hopkinson 21 22 



Difflugia lucida Penard 1902 14 21 22 25 247 

Difflugia lacustris Jung 22 25 

Difflugia lata Jung 21 22 

Difflugia linearis (Penard) Gauthier-Lievre and Thomas 22 25 

Difflugia minuta Rampi 21 22 

Difflugia paulii Ogden 21 22 

Difflugia penardi Hopkinson 21 22 244 

Difflugia pulex Penard 14 22 25 247 

Difflugia pulex var. cuneata Playfair 22 



Difflugia pyriformis Perty 244 319 

Difflugia tuberculata Playfair 24 

Difflugia tuberculata var. sphaerica Playfair 22 

Hyalospheniidae 

Heleopera petricola Leidy 22 24 319 

Heleopera petricola var amethystea Penard 22 25 

Heleopera sylvatica Penard 22 

Nebela bohemica Taranek 14 22 

Nebela collaris Leidy 22 25 260 244 258 319 

Nebela dentistoma Penard 14 22 25 



Nebela dentistoma var. oblonga Gaut.-L and Thomas 21 22 

Nebela flabellum Leidy 22 

Nebela galeata Penard 22 

Nebela lageniformis Penard 22 25 244 

Nebela militaris Penard 22 24 25 

Nebela minor Penard 22 24 

Nebela nobilis Deflandre 22 25 

Nebela parvula Penard 14 

Nebela penardiana Deflandre 21 22 25 



Nebela tincta (Leidy) Awerintzew 14 21 22 24 25 258 420 421 

Nebela tincta f. stenostoma Jung 22 

Nebela tincta var. rotunda Penard 22 

Nebela tubulosa Penard 21 22 

Nebela wailesi Deflandre 21 22 

Lesquereusiidae 

Netzelia labeosa Beyens and Chardez 23 

Microcoryciidae 

Microchlamys patella (Claparede and Lachmann) Cockerell 

Microchlamys patella var. arctica Penard 22 25 

Nebelidae 



Schönbornia smithi Beyens and Chardez 23 

Paraquadrulidae 

Paraquadrula discoides (Penard) 260 

Paraquadrula globulosa Ferlan 21 22 

Paraquadrula irregularis Deflandre 21 22 25 

Paraquadrula madarica Valkanov 21 22 

Paraquadrula ogdeni Byens and Chardez 23 

Paraquadrula rotunda Schönborn 22 260 

Phryganelidae 

Phryganella acropodia Hopkinson 14 22 24 25 244 258 260 

Phryganella acropodia var. penardi Decloitre 14 22 260 



Phryganella microps Valkanov 14 

Phryganella nidilus Penard 22 

Phryganella paradoxa var. alta Bonnet and Thomas 260 

Phryganella paradoxa Penard 22 

Plagiopyxidae 

Plagiopyxis bathystoma Bonnet 22 

Plagiopyxis callida Penard 14 21 22 25 

Plagiopyxis callida var. grandis Thomas 22 244 

Plagiopyxis declivis Thomas 14 22 

Plagiopyxis labiata Penard 14 21 22 244 



Plagiopyxis minuta Bonnet 14 260 

Plagiopyxis oblonga (Bonnet and Thomas) Bonnet 14 

Plagiopyxis penardi Thomas 22 25 

Trigonopyxidae 

Cyclopyxis ambigua Bonnet and Thomas 22 

Cyclopyxis ampulla Leidy 

Cyclopyxis arcelloides var. gibbosa van Oye 22 

Cyclopyxis arcelloides var. minima van Oye 22 

Cyclopyxis eurystoma Deflandre 22 

Cyclopyxis eurystoma var. stenostoma Decloitre 22 



Cyclopyxis kahli Deflandre 

Cyclopyxis leclerqi Beyens and Chardez 21 25 

Cyclopyxis perlucidus Beyens and Chardez 

Cyclopyxis plana var. microstoma Schönborn 14 260 

Cyclopyxis puteus Thomas 22 

Trigonopyxis arcula Penard 14 22 420 421 

Vahlkampfiidae 

Naegleria polaris De Jonckheere 2006 386 

Naegleria neopolaris De Jonckheere 2006 386 

Naegleria arctica De Jonckheere 2006 386 



Naegleria spitzbergensis De Jonckheere 2009 386 


Ciliophora 


Kinetofragminophorea 

Colpodida 

_ 

_ 

Cyrtolophosis mucicola Stokes 1885 511 

Platyophrya vorax Kahl 1926 511 

Rostrophryides sp. nov. 511 

Colpodidae 

Colpoda cucullus (Müller 1773) 319 

Cyrtophorida 

_ 

_ 

Odontochlamys alpestris Foissner 1981 511 



Chilodonella uncinata (Ehrenberg 1835) 511 

Nassulida 

_ 

_ 

Furgasonia cf. trichocystis 511 

Kreyella minuta Foissner 1979 511 

Leptopharynx costatus Mermod 1914 511 

Pleurostomatida 

_ 

_ 

Litonotus lamella (Müller 1773) 511 

Askenasia sp. 440 

Prostomatida 

_ 

_ 



Dileptus margaritifer (Ehrenberg 1833) 511 

Fuscheria cf. nodosa 511 

Phialina vertens (Stokes, 1885) 511 

Protospathidium sp. 511 

Trachelophyllum apiculatum (Perty 1852) 511 

Lacrymaria filiformis 511 

Monodinium sp. 469 

Urotrichidae 

Urotricha agilis (Stokes 1886) 511 

Urotricha cf. armata 511 


Suctorida 

_ 

_ 


Sphaerophrya terricola Foissner 1986 511 

Synhymeniida 

_ 

_ 

Nassulopsis cf. elegans 511 



Oligohymenophorea 

Hymenostomatida 

_ 

_ 

Frontonia acuminata (Ehrenberg 1833) 511 

Frontonia angusta 511 

Tetrahymena rostrata 511 

Cinetochilidae 

Cinetochilum margaritaceum (Perty 1852) 511 

Oxytrichidae 

Paramecium arcticum Doroszewski 511 

Paramecium aurelia complex 511 

Peritrichida 

_ 


_ 


Opercularia coarctata Claparede & Lachmann 1854 511 

Pseudovorticella monilata (Tatem 1870) 511 

Mobilidae 

Trichodina tentaculata 511 

Sessilidea 

Ophrydium versatile 511 

Vorticella aquadulcis complex 511 

Vorticella infusionum complex 511 

Vorticella microstoma 511 

Scuticociliatida 

_ 

_ 



Cyclidium glaucoma Müller 1773 511 

Sathrophilus muscorum (Kahl 1931) 511 

Protocyclidium muscicola (Kahl 1931) 511 



Polyhymenophorea 

Heterotrichida 

_ 

_ 

Blepharisma hyalinum Perty 1849 511 

Bryometopus pseudochilodon Kahl 1932 511 

Climacostomum virens (Ehrenberg 1838) 511 

Hypotrichida 

_ 

_ 

Aspidisca cicada (Müller 1786) 511 

Oxytricha balladyna Song & Wilbert 1989 511 

Oxytricha setigera Stokes 1891 511 



Oxytricha similis Engelmann 1862 511 

Sterkiella histriomuscorum Foissner, Blatterer, Berger & 

Kohmann 1991 

Stichotricha aculeata Wrzesniowski 1866 511 

Stylonychia mytilus complex 511 

Uroleptus gallina (Müller 1786) 511 

Oxytrichidae 

Stylonychia pustulata (Müller 1786) 

Oligotrichida 

_ 

_ 

Halteria grandinella (Müller 1773) 440 

Limnostrombidium viride (Stein 1859) 440 

511 



Rimostrombidium sp. 

Stombidium sp 440 469 


Rhizopoda 

Lobosea 

Arcellinida 

_ 

_ 


Arcellidae 

Arcella cinctus 420 421 

Centropyxidae 

Cyclopyxis arcelloides van Oye 420 421 


Apicomplexa 

Sporozoea 

Coccidia 

_ 

_ 


Sporozoea 

Sarcocystis sp. 

Toxoplasma gondii Nicolle & Manceaux 1909 530 531 544 546 547 549 550 

Isospora plectrophenaxia Dolnik and Loonen 2007 517 


Rotifera 

Digononta 

Bdelloidea 

_ 

_ 


Adinetidae 

Adineta barabata Janson 1893 46 305 319 548 

Adineta gracilis Janson 1893 46 305 319 548 

Adineta vaga (Davis 1873) 46 305 319 548 

Adineta vaga minor Bryce 1893 548 

Habrotrochidae 

Habrotrocha angusticollis (Murray 1905) 319 548 

Habrotrocha aspera (Bryce 1892) 

Callidina aspera Bryce 

319 548 



Habrotrocha bidens (Gosse 1851) 

Habrotrocha milnei Bryce 

Habrotrocha constricta (Dujardin 1841) 

Callidina constricta Dujardin 

Habrotrocha elegans (Milne 1886) 

Callidina venustra Bryce 

45 46 319 548 

45 46 305 319 548 

319 548 

Habrotrocha insignis Bryce 1915 45 46 305 319 548 

Habrotrocha lata lata (Bryce 1892) 

Callidina lata Bryce 

Habrotrocha ligula ligula Bryce 1913 548 

Habrotrocha novemdens De Koning 1947 548 

Habrotrocha pulchra (Murray 1905) 548 

319 548 



Habrotrocha pusilla (Bryce 1893) 

Callidina pusilla Bryce 

Habrotrocha pussila textrix (Bryce 1897) 319 548 

Habrotrocha roeperi (Milne 1889) 548 

Habrotrocha rosa Donner 1949 548 

Philodinidae 

Ceratrocha cornigera (Bryce 1893) 46 319 548 

Didymodactylos carnosus Milne 1916 548 

Macrotrachela aculeata Milne 1886 46 319 548 

Macrotrachela concinna (Bryce 1912) 46 319 548 

Macrotrachela ehrenbergii (Janson 1893) 46 305 319 548 

319 



Macrotrachela insolita de Koning 1947 87 548 

Macrotrachela habita (Bryce 1894) 

Callidina habita Bryce 

Macrotrachela kakkosoma (Schulte 1954) 548 

46 305 319 548 

Macrotrachela multispinosa multispinosa Thomson 1892 305 319 548 

Macrotrachela musculosa (Milne 1886) 

Callidina musculosa Milne 

Macrotrachela nana nana (Bryce 1912) 548 

Macrotrachela papillosa (Thomson 1892) 46 319 548 

Macrotrachela plicata hirundinella (Murray 1905) 548 

191 193 194 242 243 319 548 



Macrotrachela plicata plicata (Bryce 1892) 

Callidina plicata Bryce Macrotrachela plicata var hirudinella 

(Murray) 

46 319 

Macrotrachela quadricornifera Milne 1886 46 319 548 

Mniobia bredensis De Koning 1947 548 

Mniobia incrassata (Murray 1905) 319 548 

Mniobia magna (Plate 1889) 548 

Mniobia russeola (Zelinka 1891) 

Callidina russeola Zelenka 

Mniobia symbiotica (Zelinka 18886) 548 

Mniobia tentans Donner 1949 548 

45 46 319 548 



Mniobia tetraodon (Ehrenberg 1848) 

Callidina tetraodon Ehrenberg 

Mniobia vargai Donner 1962 548 

Mniobia variabilis Donner 1949 548 

Philodinia acuticornis Murray 1902 548 

Philodinia acuticornis odiosa Milne 1916 

Phiolodinia acuticornis Murray 

319 548 

46 87 319 548 

Philodinia brevipes Murray 1902 46 319 548 

Philodina erhthrophthama Ehrenberg 1832 319 

Philodina flaviceps Bryce 1906 548 

Philodinia nemoralis Bryce 1903 46 319 548 



Philodinia proterva Milne 1916 548 

Philodinia rugosa rugosa Bryce 1903 319 548 

Pleuretra alpium (Ehrenberg 1853) 

Callidina alpium Ehrenberg 

46 319 548 

Pleuretra brycei (Weber 1898) 45 46 305 319 548 

Pleuretra hystrix Bartos 1950 548 

Rotaria citrina Ehrenberg 1838 319 

Rotaria rotatoria (Pallas 1766) 

Rotaria vulgaris Schrank Rotaria rotatoria (Pallas) 

Rotaria sordida sordida (Western 1893) 

Rotifer sordida (Western) 

548 

305 319 548 



Rotaria tardigrada (Ehrenberg 1832) 

Rotifer tardigradus Ehrenberg Rotifer tardus Ehrenberg 

Scepantotrocha sp. 548 

319 548 


Monogononta 

Collothecacea 

_ 

_ 


Collothecidae 

Collotheca campanulata (Dobie 1849) 85 90 

Collotheca libera Zacharias 1894 316 

Collotheca ornata cornuta (Dobie 1842) 84 85 172 

Collotheca ornata ornuta (Ehrenberg 1832) 85 88 171 

Flosculariacea 

_ 

_ 

Filiniidae 

Filinia terminalis (Plate 1886) 246 

Flosculariidae 


Ploimida 

_ 

_ 


Ptygura melicerta (Ehrenberg 1832) 319 

Brachionidae 

Kellicottia longispina (Kellicott 1879) 

? Notholca longispina 

Keratella cochlearis (Gosse 1851) 9 87 317 

9 173 276 316 317 

Keratella hiemalis (Carlin 1943) 9 10 90 173 246 316 317 559 

Keratella quadrata (Müller 1786) 

Anuraea aculeata 

Keratella testudo (Ehrenberg 1832) 85 

7 8 9 10 319 

Notholca acuminata (Ehrenberg 1832) 9 10 85 319 



Notholca acuminata var. extensa Olofsson 1918 85 

Notholca foliacea (Ehrenberg 1838) 

Argonotholca foliacea (Ehrenberg) Notholca foliacea var. latistyla 

Olofsson 

Notholca labis Gosse 1887 83 

7 9 83 85 90 171 172 246 316 

Notholca latistyla (Olofsson 1918) 9 83 85 90 173 

Notholca latistyla f. amreni De Smet and Bafort 1990 84 85 

Notholca marina (Focke 1991) 85 

Notholca striata (Müller) 

Notholca striata var. bipalium 

(Müller) 

Notholca striata f. extensa 

9 10 319 

Notholca squamala (Müller 1786) 9 10 84 85 88 90 171 172 317 559 

Colurellidae 



Colurella adriatica Ehrenberg 1831 

Colurella amblytelus (Gosse 1851) 319 

Colurella colurus (Ehrenberg 1830) 83 85 89 171 172 319 

Colurella geophila Donner 1981 83 

Colurella hindenburgi Steinecke 1917 83 85 90 317 

Colurella obtusa (Gosse 1886) 83 85 319 

Colurella uncinata (Müller 1773) 83 85 90 

Lepadella acuminata (Ehrenberg 1834) 

Metopidia acuminata Ehrenberg 

Lepadella minuta (Montet 1918) 83 85 

83 85 90 319 



Lepadella ovalis (Müller 1786) 

Metopidia lapadella 

Lepadella patella (Müller 1786) 

Metopidia oblonga 

Lepadella quadricarinata (Stenroos 1898) 

Metopidia quadricarinata 

Lepadella triba Myers 1934 85 

83 85 173 172 319 

46 83 84 85 88 90 171 172 305 

83 85 319 

Lepadella triptera Ehrenberg 1830 83 316 

Lophocaris oxysternon (Goose 1851) 319 

Dicranophoridae 

Albertia naidis Bousfield 1886 83 

Dicranophorus forcipatus (Müller 1786) 83 85 



Encentrum bidentatum (Lie-Pettersen 1906) 86 

Encentrum dieteri De Smet 1995 86 

Encentrum felis (Müller 1773) 

Encentrum ferox (Western) nom. dub 

Encentrum ferox (Western) nom. dub. 

Encentrum grandi Haring and Myers 1928 

Encentrum grandi (Western) nom. 

dub 

Encentrum marinum (Dujardin 1841) 84 85 

Encentrum mucronatum Wulfert 1936 84 87 88 

Encentrum murrayi Bryce 1922 46 319 

Encentrum mustela (Milne 1885) 85 

319 

06 June 2013 Page 53 of 185 

62


Encentrum permolle (Gosse 1886) 

Dicranophorus permollis permollis 

(Gosse) 

Encentrum uncinatum (Milne 1886) 

Diglena permollis Gosse 

Encentrum uncinatus (Milne) Dicranophorus uncinatus (Milne) 

Itura aurita aurita (Ehrenberg 1930) 

Itura aurita var. intermedia (Wulfert) 

87 308 

Epiphanidae 

Rhinoglena frontalis Ehrenberg 1858 83 85 

84 88 90 171 172 

Euchlanis deflexa Gosse 1851 83 85 319 

Euchlanis dilatata Ehrenberg 1832 9 83 85 171 172 319 

Euchlanis lyra Hudson 1886 89 

Euchlanis meneta Myers 1930 

Euchlanis oropha Gosse 

85 

83 85 316 319 



Catyphna brevis Murray 1913 319 

Catyphna rotundata Olofsson 1918 319 

Lecanidae 

Lecane bulla (Goose 1851) 

Monostyla bulla Gosse 

46 319 

Lecane closterocerca (Schmarda 1859) 83 87 

Lecane cornuta (Müller 1786) 

Monostyla cornuta (Müller) 

Lecane flexilis (Gosse 1886) 

Lecane brevis (Murray) 

Lecane ligona (Dunlop 1901) 317 

Lecane lunaris (Ehrenberg 1832) 

Monostyla lunaris (Ehrenberg) 

305 319 

83 319 

46 83 171 172 305 319 



Lecane pideis (Harring and Myers 1926) 

Lecane piepelsi de Smet and Bafort 

Lecane robertsonae Segers 1993 

Lecane rotundata (Olofsson) 

Lecane rotundata (Oloffson 1918) 

Lindiidae 

Lindia cf. torulosa Dujardin 1841 83 

85 171 172 

Mytilinidae 

Lophocharis oxysternon (Gosse 1851) 83 85 

Mytilina bicarinata (Perty 1850) 

62 83 171 319 

Mytilina mucronata (Müller 1773) 83 85 90 171 172 319 

Mytilina ventralis var. brevispina (Ehrenberg 1832) 

Mytilina brevispina Ehrenberg 

Notommatidae 

46 171 172 305 319 



Cephalodella biungulata Wulfert 1937 85 

Cephalodella catellina (O.F. Müller 1786) 83 85 90 171 172 

Cephalodella evabroedi De Smet 1988 83 85 

Cephalodella glandulosa Koch-Althaus 1962 85 

Cephalodella gibba (Ehrenberg 1832) 

Diaschiza gibba (Ehrenberg) 

Cephalodella gracilis (Ehrenberg 1832) 

Diaschiza gracilis (Ehrenberg) 

Cephalodella hoodi (Goose 1896) 83 85 

Cephalodella intuta Myers 1924 85 90 

Cephalodella megalocephala (Glascott 1893) 85 

83 84 85 88 90 171 172 319 

319 



Cephalodella misgurnus Wulfert 1937 84 88 

Cephalodella rotunda Wulfert 1937 85 

Cephalodella sterea (Gosse 1887) 84 88 

Cephalodella ventripes (Dixon-Nuttall 1901) 171 172 

Cephalodella ventripes var. angustior Donner 1949 83 85 

Diaschiza spp. 

? Cephalodella sp. 

Eosphora najas Ehrenberg 1830 83 85 

Notommata cyrtopus Gosse 1886 83 

Notommata glyphura Wulfert 1935 85 

319 



Resticula nyssa Harring and Myers 1924 83 85 

Proalidae 

Bryceella stylata (Milne 1886) 

Squatinella stylata (Milne) Stephanops stylatus Milne 

Bryceella tenella (Bryce 1897) 

Squatinella tenella (Bryce) Stepanops tenullus Bryce 

Proales decipiens (Ehrenberg 1832) 

Furcularia decipiens Ehrenberg 

Proales fallaciosa Wulfert 1937 85 

Proales sordida Gosse 1886 319 

Wulfertia ornata Donner 1943 85 

Scaridiidae 

Scaridium longicaudum (Müller 1786) 319 

Synchaetidae 

85 319 

319 

319 



Polyarthra dolichoptera (Idelson 1925) 7 9 10 83 85 90 173 246 316 317 559 

Polyarthra dolichoptera f. aptera (Hood 1895) 85 

Polyarthra dolichoptera f. proloba (Albertova 1960) 85 

Polyarthra vulgaris Carlin 1943 319 

Synchaeta lakowitziana Ehrenberg 1832 9 10 

Synchaeta lakowitziana arctica De Smet 1988 883 317 

Synchaeta littoralis Rousselet 1902 9 10 

Synchaeta truncata von Hofsten 9 10 

Trichoceridae 

Trichocerca bidens (Lucks 1912) 

Diurella bidens Lucks 

90 319 



Trichocerca. cavia (Gosse 1886) 85 171 172 

Trichocerca cristata Harring 319 

Trichocerca intermedia (Stenroos 1898) 85 

Trichocerca longistyla (Olofsson 1918) 85 

Trichocerca rattus (Müller 1776) 

Rattulus carinatus 

83 85 319 

Trichocerca relicta (Donner 1950) 83 85 

Trichocerca uncinata (Voigt 1902) 

Diurella uncinata (Voigt) 

83 85 90 319 

Trichocerca weberi (Jennings 1903) 83 85 90 171 172 

Trichotria truncata (Whitelegge) 191 193 242 



Trichotria pocillum (Müller 1776) 83 


Gastrotricha 

_ 


Chaetonotoida 

_ 

_ 

Chaetonotidae 

Chaetonotus sp. 88 


Nematoda 

Adenophorea 

Enoplia 

Stichosomida 

_ 

_ 


Capillariidae 

Capillaria sp 506 

Philometridae 

Philonema oncorhynchi (Kuitenen Ekbaum 1933) 184 

Trichinellidae 

Trichinella spiralis (Owen 1935) 

Trichinella nelsnoni 

Trichinella nativa (Britov and Boev 1972) 180 

36 189 190 207 


Penetrantia 

Dorylaimida 

_ 

_ 


Dorylaimidae S.L. 

Enchodelus analatus Thorne 1939 215 

Enchodelus conicaudatus Thorne 1939 215 

Enchodelus cf. macrodoroides (Steiner 1914) 215 

Enchodelus macrodorus Thorne 1939 215 

Enchodelus parvus Loof 1971 215 

Epidorylaimus agilis (de Man 1880) Andrassy 1986 

Eudorylaimus agilis Loof Dorylaimus lugdunensis apud 

Steiner nec. de Man 

215 



Epidorylaimus lugdunensis (de Man 1880) 

Eudorylaimus lugdunensis 

Andrassy, 1959 

Eudorylaimus alleni Brzeski1962 215 

Eudorylaimus carteri (Bastian 1865) 

Dorylaimus carteri Bastian 

Eudorylaimus centrocercus (de Man 1880) 

Dorylaimus centrocercus de Man 

Eudorylaimus circulifer (Loof 1961) 215 

215 

191 193 194 243 

Eudorylaimus maksymovi (Altherr 1963) 191 193 215 

Eudorylaimus megodon Loof 1971 215 

Eudorylaimus parvus (de Man 1880) 215 


6


Eudorylaimus subjunctus Loof 1971 215 

Eudorylaimus vanrosseni Loof 1971 215 

Longidorella magna Loof 1971 215 

Mesodorylaimus sp. 171 

Mononchidae 

Prionchulis muscorum (Dujardin 1845) 191 193 

Qudsianematidae 

Labronema sp. 191 193 

Tylencholaimidae 

Tylencholaimus proximus Thorne 1939 215 

Enoplida 

_ 

Tylencholaimus teres Thorne 1939 215 


_ 


Alaimidae 

Alaimus arcuatus Thorne 1939 215 

Alaimus depressus Loof 1971 215 

Alaimus parvus Thorne 1939 215 

Prismatolaimidae 

Prismatolaimus dolichurus de Man 1880 6 215 

Prismatolaimus intermedius (Buetschli1873) 6 215 

Prismatolaimus primitivus Loof 1971 215 

Prismatolaimus stenolaimoides Loof 1971 215 

Tobrilidae 

Tobrilus sp. 171 191 193 

Tripylidae 



Tripyla cf. infia Brzeski et Winiszewska-Slipinska 1903 171 

Tripyla glomerans Bastian 1865 

Tripyla papillata Butschli 

215 


Secernentia 

Ascaridida 

_ 

_ 


Anisakidae 

Anisakis simplex (Rudolphi 1802) 468 

Contracaecum osculatum (Rudolphi 1801) Baylis 1920 468 

Hysterothylacium aduncum (Rudolphi 1802) 506 

Rhabditida 

_ 

_ 

Bunonematidae 

Bunonema reticulatum Richters 1905 215 

Cephalobidae 

Acrobeles ciliatus von Linstow 1877 215 



Acrobeloides bütschlii Gadea 1954 

Cephalobus bütschlii de Man 

Acrobeloides enoploides Loof 1971 215 

Acrobeloides tricornis (Thorne 1925) 38 215 

Cervidellus serratus (Thorne 1925) 215 

Cervidellus spitzbergensis Bostrøm 1987 37 

Chiloplacus saccatus Loof 1971 38 215 

Eucephalobus arcticus Loof 1971 38 215 

Heterocephalobus elongatus (de Man 1880) 

Cephalobus elongatus de Man 

Stegelleta similis (Thorne 1925) 

Stegelleta mucronata 

228 

215 


6 215


Panagrolaimidae 

Panagrolaimus papillosus Loof 1971 215 

Teratocephalidae 

Euteratocephalus crassidens Andrassy 1958 

Metateratocephalus crassidens (de 

Man) 

Teratocephalus costatus Andrassy 1958 39 

Teratocephalus decarinus Anderson 1969 215 

6 39 215 

Teratocephalus lirellus Anderson 1969 39 215 

Teratocephalus terrestris (Butschli1873) 6 

Spirurida 

_ 

_ 

Acuriidae 

Paracuaria adunca (Creplin 1846) Anderson and Wong 1981 468 506 



Stegaphorus stellae-polaris (Parona 1901) 360 468 506 

Strongylida 

_ 

_ 

Trichostrongylidea 

Marshallagia marshalli m occidentalis (Ransom, 1907) 

Grosspiculaqia occidentalis 

(Ransom 1907) 

54 131 132 134 385 522 586 

Marshallagia marshalli (Ransom 1907) 5 48 50 53 54 131 132 133 134 369 384 385 430 

431 432 439 489 490 522 586 594 606 

Nematodirus helvetianus May 1920 5 133 

Ostertagia arctica Mitzkewitzsch 1929 5 133 134 

Ostertagia grühneri Skrjabin 1929 2 3 5 48 50 53 54 131 132 133 346 369 383 

384 430 431 432 439 488 489 490 383 522 586 

Ostertagia occidentalis Ransom 1907 5 48 50 53 


586 

489


Paracuaria adunca (Creplin 1846) 257 

Skrjabinagia arctica (Mitzkevich 1929) 

?Skrjabinagia lyrata (Sjøberg 1926). 

48 49 53 54 131 132 134 522 

Teladorsagia circumcincta (Stadelmann 1894) 48 49 53 54 131 132 133 134 522 

Teladorsaga davitiani Andreeva and Satubaldini1984 54 131 134 522 

Teladorsagia trifurcata 522 

Tylenchida 

_ 

_ 

Aphelenchinae 

Aphelenchus nivalis Aurivillius 1883 306 228 

Criconematidae 

Criconemoides annulatus Taylor 1936 

Criconemoides hemispaericaudatus 

Wu 

215 



Criconema sp. 215 

Hoplolaimidae 

Helicotylenchus spitsbergensis Loof 1971 215 

Pratylenchidae 

Pratylenchoides crenicauda Winslow 1958 215 

Pratylenchoides magnicauda (Thorne 1935) 

Tylenchorhynchus magnicauda 

Filipijev 

Tylenchidae 

Basiria dolichura Loof 1971 215 

Coslenchus costatus (de Man 1921) 

Tylenchus costatus de Man 

Filenchus thornei (Andrassy 1954) 

Tylenchus thornei Andrassy 

215 

215 

215 



Geocenamus arcticus (Mulvey 1969) 

Tylenchorhynchus arcticus Mulvey 

Geocenamus loofi (Siddiqi1979) 

Tetylenchus joctus Thorne apud Loof Merlinius loofi Siddiqi 

Geocenamus microdorus (Geraert 1966) 

Tylenchorhynchus microdorus 

Geraert 

Lelenchus leptosoma (de Man 1880) 

Tylenchus leptosoma de Man 

Malenchus bryophilus (Steiner 1914) 

Tylenchus bryophilus Steiner 

Nagelus leptus (Allen 1955) 

Tylenchorhynchus leptus Allen 

Tylenchus davainei Bastien 1865 

Anguillulina davainei (Bastian) 

215 

215 274 

215 

215 

215 

215 


6 215

Torquentia 

Araeolaimida 

_ 

_ 


Plectidae 

Anaplectus cf. grandepapillatus (Ditlevsen 1928) 171 

Anaplectus granulosus de Coninck and Shürmans Stekhoven 

1933 

Plectus granulosus Bastian 

Anaplectus porosus Allen and Noffsinger 1968 191 193 194 215 

Ceratoplectus communis (Bütschli1873) 

Plectus communis Bütschli 

Ereptonema arcticum Loof 1971 38 215 

Plectus acuminatus Bastian 1865 215 

Plectus cf armatus Butschli1873 215 

228 



Plectus assimilis Butschli1873 215 

Plectus aquatilis Andrassy1984 171 

Plectus cirratus Bastian 1865 6 191 193 194 228 

Plectus cornus Maggenti1961 215 

Plectus geophilus de Man 1880 215 

Plectus inquirendus Andrassy 1958 215 

Plectus longicaudatus Butschli1873 215 

Plectus cf. opisthocirculus Andrassy 1952 171 

Plectus parietinus Bastian 1865 171 191 193 194 215 243 



Plectus parvus Bastian 1865 215 

Plectus rhizophilus de Man 1880 215 

Rhabdolaimidae 

Rhabdolaimus terrestris de Man 1880 215 

Chromadorida 

_ 

_ 

Achromadoridae 

Achromadora cf. semiarmata Altherr 1952 215 

Achromadora tenax (de Man 1876) 171 215 

Ethmolaimidae 

Ethmolaimus sp. 

Odontolaimus 

Odontolaimus chlorurus de Man 1880 215 

Monohysterida 


_ 

_ 


Monhysteridae 

Eumonhystera dispar (Bastian 1865) 

Monohystera cf. dispar Bastian 

Eumonhystera filiformis (Bastian 1865) 

Monohystera filiformis Butschli 

Eumonhystera vulgaris (de Man 1880) 

Monohystera vulgaris de Man 

Geomonohystera villosa (Butschli1873) 

Monohystera villosa Butschli 

215 

171 215 

Monohystera stagnalis Bastian 1865 171 215 

215 



Acanthoceophala 

Palaecanthocephala 

Polymorphida 

_ 

_ 

Polymorphidae 

Corynosoma strumosum (Rudolphi 1802) Luhe 1904 468 506 

Notocotylus sp. 506 

Acantocephalan sp. 545 


Platyhelminthes 

Cestoda 


Cyclophyllidae 

_ 

_ 

Anoplocephalidae 

Moniezia benedini Moniez 1872 48 49 50 51 131 

Dilepididae 

Anomotaenia campylacantha (Krabbe 1869) 360 

Anomotaenia larina (Krabbe 1869) 

Alcataenia larina 

468 506 

Anomotaenia micacantha (Krabbe 1869) 468 506 

Alcataenia (Rissotaenia) dominicana (Raillet and Henry 1912) 468 506 

Arctotaenia tetrabothriodes 506 



Paricteropaenia porosa (Rudolphi 1810) 468 

Hymenolepididae 

Alopparakis larina Furhmann 1921 

Microsomacanthus ductilus 506 

Microsomacanthus microsoma (Creplin, 1829) Lopez-Neyra, 

1942 

Microsomacanthus diorchis (Fuhrmann, 1913). 506 

Taeniidae 

Echinococcus multilocularis Leuckart 1863 399 438 515 545 581 582 588 

Taenia crassiceps 545 

Taenia ovis krabbei (Moniez 1879) Verster 1969 

Cysticercus tarandi 

506 

48 50 50 131 545 560 



Taenia polyacantha 545 

Tetrabothrius minor (Loennberg, 1893) 506 

Tetrabothriidae 

Tetrabothrius erostris (Loennberg 1889) 468 506 

Proteocephalidea 

_ 

_ 

Proteocephalidae 

Proteocephalus exiguus Weinland 1858 184 396 

Pseudophyllidea 

_ 

Diphyllobothriidae 

Diphyllobothriidae 

Diphyllobothrium ditremum (Creplin 1825) 184 195 396 491 

Triaenophoridae 

Eubothrium crassum (Bloch 1779) 183 



Eubothrium salvelini Schrank 1790 183 184 277 396 


Trematoda 


Opisthorchiida 

Opisthorchiata 

_ 

Heterophyidae 

Cryptocotyle lingua (Creplin 1825) 257 468 506 

Plagiorchiida 

Brachylaimata 

_ 

Microphallidae 

Microphallus pygmaeus (Levinsen,1881) 506 

Microphallus pseudopygmaeus Galaktionov,1980 506 

Microphallus triangulatus 506 

Strigeata 

Plagiorchiata 

_ 

Gymnophallidae 



Gymnophallus somateria (Levinsen,1881) 506 

Gymnophallus choledochus Odhner,1900 506 


Annelida 

Oligochaeta 

Haplotaxida 

_ 

_ 


Enchytraeidae 

Bryodrilus diverticulatus Cernosvitov 1929 

Bryodrilus ehlersi glandulosa 

30 95 354 351 

Bryodrilus ehlersi Ude 1892 28 30 95 236 237 238 283 

Bryodrilus librus (Neilsen and Christensen 1959) 

Marionina libra Neilsen and 

Christensen 1959 

95 236 238 595 

Bryodrilus parvus Nurminen 1970 28 30 95 283 351 

Cernosvitoviella immota (Knoller 1935) 30 95 236 237 238 

Cernosvitoviella cf. Pusilla 



Cognettia glandulosa (Michaelsen 1888) 580 595 

Cognettia sphagnetorum (Vejdovsky 1877) 95 237 238 

Enchytraeus albidus Henle 1837 95 236 238 289 290 291 305 306 595 

Enchytaeus dichaetus Schmelz & Collado 2012 580 

Enchytraeus kincaidi Eisen 1904 

Enchytraeus liefdeensis Stephenson 

Fridericia bulboides Neilsen and Christensen 1959 95 236 238 

30 95 236 237 238 290 291 306 

Fridericia bulbosa (Rosa 1887) 95 236 237 238 266 

Fridericia leydigi (Vejdovsky 1877) 95 236 

Henlea heleotropha Stephenson 1922 augm. Nielsen and 

Christensen 1959 

Henlea heleotrophus Stephenson 



Henlea glandulifera Nurminen 1970 

Henlea similis 

Henlea nasuta (Eisen 1878) 95 595 

Henlea perpusilla Friend 1911 

Henlea brucei Stephenson 

95 237 238 354 351 

Henlea similis Nielsen and Christensen 1959 28 30 236 237 238 302 

28 30 95 236 237 238 266 283 350 351 373 363 595 

Henlea ventriculosa (d’Udekem 1954) 28 30 236 237 238 302 351 595 

Lumbricillus arenarius (Michaelsen 1988) 

Lumbricillus magdalenae Nurminen 

Lumbricillus bülowi Nielsen and Christiensen 1959 95 

Lumbricillus eltoni (Stephenson 1924) 

Enchytraeus eltoni Stephenson 

95 236 238 

95 236 290 



Lumbricillus muscicolus (Stephenson 1924) 

Enchytraeus muscicolus Stephenson 

Lumbricillus neilseni Nurminen 1965 

as Lumbricillus bülowi Nielsen and 

Christiensen 1959 ref. 30 

Lumbricillus pagenstecheri (Ratzel 1869) 

as Lumbricellus aegialites 

Stepheson 1922 

as Lumbricellus nervosus Ude 1902 

as Lumbricellus necrophagus 

Stephenson 1922 

95 236 291 

95 236 237 238 

Lumbricillus reynoldsoni Backlund 1948 95 236 238 

Lumbricillus rivalis Levinsen 1890 

Lumbricillus semifuscus (Claparede 1861) augm. Stephenson 

1911 

Lumbricillus tuba Stephenson 1911 95 

Marionina aporus (Stephenson 1925) 

Enchytraeus aporus Stephenson 

95 236 238 289 290 291 305 306 

95 236 291 



Marionina argentea (Michaelsen 1889) 28 30 95 236 237 238 266 

Marionina crymodes (Stephenson 1922) 

Enchytreaus crymodes Stephenson 

95 236 289 305 306 

Mesenchytraeus argentatus Nurminen 1973 28 30 95 283 351 

Mesenchytraeus eltoni Stephenson 1925 95 236 291 306 

Mesenchytraeus flavus (Levinsen 1884) 28 30 95 237 238 283 

Mesenchytraeus melanocephalus 

Mesenchytraeus sp. n. 

Lumbricidae 

Dendrodrilus rubidis (Savigny 1826) 580 595 

Dendrobaena hortensis (Michaelsen 1890) 580 595 


Tardigrada 

Eutardigrada 

Apochela 

_ 

_ 


Milnesiidae 

Milnesium asiaticum Tumanov 2006 576 593 

Milnesium eurystomum Mauicci 1991 576 593 

Milnesium tardigradum Doyère 1840 78 221 222 254 319 336 593 

Parachela 

_ 

_ 

Hypsibiidae 

Acutuncus antarcticus (Richters 1904) 

Hypsibius antarcticus (Richters 

1905) 

593 



Borealibius zetlandicus (Murray 1907) 

Hypsibius (Hypsibius) zetlandicus 

(J. Murray) 

Diphascon (Diphascon) stappersi Richters 1911 

Hypsibius stappersi (Richters) 

Diphascon (Diphascon) alpinum Murray 1906 

Hypsibius alpinus (Murray) 

Diphascon (Diphascon) chilenense Plate 1888 

Hypsibius zetlandicus (J.Murray 

1907) 

Diphascon chilenense (Plate) Hypsibius chilenensis Plate 

Diphascon (Diphascon) oculatum oculatum Murray 1906 

Diphascon oculatum Murray 

Diphascon (Diphascon) pingue pingue (Marcus 1936) 

Diphascon pingue (Marcus) 

Diphascon (Diphascon) recamieri Richters 1911 

Hypsibius recamieri (Richters) Diphascon recamieri Richters 

78 171 221 222 593 

222 254 319 329 336 593 

90 171 222 233 251 328 336 593 

221 222 233 254 336 593 

78 222 593 

78 221 222 329 330 576 593 

78 87 171 221 222 251 330 336 576 593 



Diphascon (Diphascon) tenue Thulin 1928 

Diphascon tenue Thulin Hypsibius tenuis (Thulin) 

Diphascon (Adropion) arduifrons Thulin 1928 

Diphascon arduifrons Thulin 

Diphascon (Adropion) behanae Dastych, 1987 593 

Diphascon (Adropion) belgicae Richters 1911 

Fujiscon belgicae (Richters) 

Hypsibius belgicae (Richters) 

Diphascon (Adropion) prorsirostre Thulin 1928 

Diphascon prorsirostre Thulin 

Diphascon (Adropion) scoticum J.Murray 1905 

F. diphasconiellum Ito 

Diphascon belgicae Richters 

Diphascon scoticum J.Murray Hypsibius scoticus (J.Murray) 

78 221 222 593 

78 222 593 

78 221 222 254 319 336 593 

221 222 593 

Doryphoribius macrodon Binda Pilato and Dastych 1980 27 78 222 593 

Hebesuncus conjungens (Thulin 1911) 

Diphascon conjugens (Thulin) 

78 90 171 222 233 251 329 330 576 593 

222 229 233 576 593 



Hypsibius biscuitiformis Bartos, 1960 593 

Hypsibius arcticus (Murray 1907) 

Macrobiotus arcticus Murray 

Hypsibius convergens (Urbanowicz 1925) 78 171 221 222 229 251 329 593 

Hypsibius dujardini (Doyère 1840) 

Macrobiotus tetradactylus Greef 

Hypsibius microps Thulin 1928 221 222 576 593 

Hypsibius pallidus Thulin 1911 77 78 222 576 593 

Isohypsibius annulatus annulatus (Murray 1905) 593 

Isohypsibius dastychi Pilato Bertolani and Binda 1982 78 222 593 

Isohypsibius coulsoni 576 593 

593 

78 89 90 171 221 222 229 233 576 593 



Isohypsibius elegans Binda and Pilato 1971 78 222 330 593 

Isohypsibius granulifer granulifer Thulin 1928 

Hypsibius granulifer (Thulin 1928) 

Isohypsibius papillifer papillifer (Murray 1905) 

Hypsibius papillifer (J.Murray) 

78 90 87 171 222 251 328 329 593 

222 328 329 330 593 

Isohypsibius papillifer bulbosus (Marcus 1928) 222 328 329 336 593 

Isohypsibius marcellinoi Binda and Pilato, 1971 593 

Isohypsibius prosostomus prosostomus Thulin 1928 

Hypsibius prosostomus (Thulin) 

Isohypsibius sattleri (Richters, 1902) 593 

Isohypsibius schaudinni (Richters 1909) 

Hypsibius schaudinni (Richters) 

78 171 221 222 593 

222 336 593 



Isohypsibius tetradactyloides (Richters, 1907) 593 

Isohypsibius tuberculatus (Plate 1888) 

Macrobiotus tuberculatus Plate Hypsibius tuberculatus Plate 

Mesocrista spitzbergensis (Richters 1903) 

Hypsibius spitzbergensis Richters Diphascon spitzbergense (Richters) 

Platicrista angustata (Murray 1905) 

Diphascon angustatum Murray Hypsibius angustatus (Murray) 

Ramazzottius cataphractus (Maucci1974) 

Hypsibius cataphractus Maucci 

Ramazzottius montivagus (Dastych 1983) 

Hypsibius montivagus Dastych 1983 

Ramazzottius oberhaeuseri (Doyère 1840) 

Macrobiotus oberhauseri (Doyère) 

Hypsibius oberhauseri Doyère 

Macrobiotus oberhauseri var. 

granulatus Richters 

222 233 336 319 593 

90 171 191 192 193 194 222 233 593 

78 221 222 233 336 576 593 

78 221 222 593 

78 221 222 329 593 

78 221 222 233 254 319 336 593 



Thulinius augusti (Murray 1907) 

Pseudobiotus augusti (Murray) Hypsibius (Isohysibius) augusti 

Murray 

Halobiotus arcturulius Crisp and Kristensen, 1983 593 

Halobiotus crispae Kristensen, 1983 593 

222 229 230 593 

Macrobiotidae 

Macrobiotus ariekammensis Weglarska 1965 78 222 593 

Macrobiotus crenulatus Richters 1904 26 221 222 233 330 576 593 

Macrobiotus echinogenitus Richters 1903 

? Macrobiotus crenulatus Richters 

Macrobiotus grandis Richters, 1911 593 

26 78 90 191 192 193 222 233 

Macrobiotus harmsworthi harmsworthi Murray 1907 78 221 222 233 254 319 330 336 576 593 



Macrobiotus harmsworthi obscurus Dastych 1985 78 192 193 222 576 593 

Macrobiotus hufelandi hufelandi C.A.S. Schultze 1833 

Macrobiotus hufelandi Schultze 

1833 

78 221 222 233 251 319 328 336 576 593 

Macrobiotus islandicus islandicus Richters 1904 78 191 192 193 221 222 233 576 593 

Macrobiotus occidentalis occidentalis Murray, 1910 593 

Minibiotus crassidens (Murray, 1907) 593 

Minibiotus intermedius (Plate 1888) 

Macrobiotus intemedius Plate 

Paramacrobiotus areolatus ( Murray 1907) 

Macrobiotus areolatus Murray 1907 

Paramacrobiotus richtersi (Murray 1911) 

Macrobiotus richtersi Murray 1911 

78 221 222 233 319 336 593 

221 222 336 593 

78 221 222 229 230 593 



Tenuibiotus voronkovi (Tumanov, 2007) 593 

Tenuibiotus willardi (Pilato, 1977) 593 

Richtersius coronifer (Richters 1903) 

Adorybiotus coronifer (Richters) Macrobiotus coronifer Richters 

Calohypsibiidae 

Calohypsibius ornatus (Richters 1900) 

Macrobiotus ornatus var 

spinosissimus Richters 

Eohypsibiidae 

Bertolanius nebulosus (Dastych 1983) 

Amphibolus nebulosus (Dastych 

1983) 

Bertolanius smreczynskii (Weglarska 1970) 

Doryphoribius smreczynskii 

Weglarska 

Bertolanius weglarskae (Dastych 1972) 

Amphibolus weglarskae (Dastych 

1972) 

Hypsibius ornatus Richters 

Amphibolus smreczynskii 

(Weglarska 1970) 

78 221 222 319 336 593 

221 222 233 254 319 336 576 593 

77 78 222 593 

90 171 191 192 193 194 222 243 593 

222 328 329 330 593 



Murrayidae 

Dactylobiotus dispar (Murray 1907) 

Macrobiotus dispar Murray 

Dactylobiotus macronyx (Dujardin 1851) 

Macrobiotus macronyx Dujardin 

Murrayon dianeae (Kristensen 1982) 

Macrobiotus dianeae Kristensen 

1982 

Murrayon hibernicus (Murray 1911) 

Macrobiotus hibernicus (Murray 

1911) 

Murrayon pullari (Murray 1907) 

Macrobiotus pullari Murray Macrobiotus pullari (Murray 1907) 

191 192 193 222 233 329 330 593 

222 234 305 336 593 

222 329 330 593 

78 222 593 

89 90 222 233 336 593 


Heterotardigrada 


Arthrotardigrada 

_ 

_ 

Echiniscidae 

Bryodelphax parvus Thulin 1928 593 

Bryodelphax parvuspolaris Kacmarek 2012 576 593 

Bryodelphax sinensis (Pilato 1974) 

Echiniscus sinensis Pilato 

78 221 222 593 

Echiniscus arctomys Ehrenberg 1853 222 233 319 336 593 

Echiniscus blumi blumi Richters 1903 78 221 222 319 336 593 

Echiniscus capillatus Ramazzotti1956 78 222 593 

Echiniscus columinis Murray, 1911 593 



Echiniscus granulatus (Doyère 1840) 78 221 222 254 593 

Echiniscus merokensis merokensis Richters 1904 78 221 222 336 576 593 

Diploechiniscus oihonnae (Richters 1903) 

Echiniscus oihonnae Richters 1903 

222 254 336 593 

Echiniscus quadrispinosus quadrispinosus Richters 1902 222 336 576 593 

Echiniscus spiculifer Schaudinn, 1901 593 

Echiniscus spinulosus (Doyère 1840) 222 336 593 

Echiniscus testudo (Doyère 1840) 78 221 222 336 576 593 

Echiniscus trisetosus Cuenot, 1932 593 

Pseudechiniscus islandicus (Richters 1904) 222 330 593 



Pseudechiniscus suillus (Ehrenberg 1853) 78 221 222 329 330 336 576 593 

Pseudechiniscus victor (Ehrenberg 1853) 78 221 222 254 336 593 

Testechiniscus spitsbergensis (Scourfield 1897) 

Echiniscus spitsbergensis Scourfield 

78 221 222 254 319 329 330 336 576 593 


Bryozoa 

Phylactolaemata 

_ 

_ 


Statoblasts. Species unknown 519 


Chelicerata 

Arachnida 

Acari 

Acariformes 


Oribatida 

Ameronothroidea 

Ameronothridae 

Ameronothrus lineatus (Thorell 1871) 

Eremaeus lineatus Thorell Scutovertex lineatus (Thorell) 

Ameronothrus nidicola Sitnikova 1975 

Probably Ameronothrus lineatus 

(Thorell) 

34 41 42 70 164 181 182 219 254 271 305 306 318 

319 342 343 363 373 442 453 480 481 482 483 484 485 

499 501 555 563 567 

56 567 

Brachychthonoidea 

Brachychthoniidae 

Eobrachychthonius borealis (Forsslund 1942) 235 442 567 

Eobrachychthonius latior (Berlese 1910) 

Eobrachychthonius grandis (Sellnick) 

Eobrachychthonius oudemansi van der Hammen 1952 235 442 567 

181 182 219 235 442 567 


453


Liochthonius alpestris (Forsslund 1958) 56 567 

Liochthonius brevis (Michael 1888) 

Brachychthonius perpusillus 

Berlese 

Liochthonius perpusillus (Berlese) 

Liochthonius laetepictus (Berlese 1910) 

Brachychthonius laetepictus Berlese 

Brachychthonius brevis (Michael) 

var. glabra 

181 318 320 442 567 595 

181 182 235 320 442 567 

Liochthonius lapponicus (Trägårdh 1910) 56 119 181 182 442 563 567 

Liochthonius muscorum Forsslund 1964 56 119 567 595 

Liochthonius neglectus Moritz 1976 457 567 

Liochthonius sellnicki (Thor 1930) 

Brachychthonius sellnicki Thor 

Liochthonius simplex 563 

181 182 219 235 318 320 445 442 567 595 



Liochthonius strenzki Forsslund 1942 457 567 

Neoliochthonius piluliferus (Forsslund 1942) 457 

Malaconothridae 

Malaconothrus monodactylus (Michael 1888) 

Malaconothrus egregius (Berlese 

1904) 

Phthiacaroidae 

Atropacarus striculus (C.L. Koch 1836) 

Steganacarus striculus (C.L. Koch 

1836) 

441 442 567 

441 442 526 

Carabodoidea 

Carabodidaae 

Carabodes marginatus (Michael, 1884) 526 567 

Tectocepheidae 

Tectocepheus alatus Berlese 1913 235 442 567 

Tectocepheus sarekensis Trägårdh 1910 181 182 442 567 



Tectocepheus velatus (Michael 1880) 56 70 181 182 219 318 320 377 387 420 421 442 563 

567 595 

Tectocepheus velatus knullei Vanek 1960 442 526 567 

Ceratozetoidea 

Ceratozetidae 

Ceratozetes spitsbergensis Thor 1934 70 181 182 272 320 442 567 

Chamobates borealis (Trägårdh 1902) 56 567 

Chamobates birulai (Kulcynski1902) 

Notaspis birulai Kulcynski 

319 567 

Chamobates cuspidatus (Mihael, 1844) 528 567 

Diapterobates notatus (Thorell 1871) 

Oribata notata Thorell 

Trichoribates notatus (Thorell) 

Sphaerozetes notatus Thorell 

Murcia notata (Thorell) 

13 25 34 56 66 69 70 121 152 155 157 164 181 

191 235 271 182 266 272 305 306 318 319 320 335 342 

363 373 377 387 411 418 420 421 442 501 526 553 555 

563 567 567 

305 



Edwardzetes edwardsii (Nicolet 1855) 181 182 320 442 567 

Iugoribates gracilis Sellnick 1944 457 567 

Mycobates parlmeliae (Michael 1884) 442 526 567 

Mycobates sarekensis (Trägårdh 1910) 

Calyptozetes sarekensis (Trägårdh) 

Mycobates tridactylus Willmann 1929 442 567 

Cytozetes sp. 563 

Oromurcia lucens (L.Koch 1879) 

Notaspis lucens L. Koch Probably Svalbardia paludicola 

Thor, 1930 

69 181 182 219 235 273 318 320 377 411 420 421 442 

501 563 567 

319 567 567 

Punctoribates punctum (C.L. Koch 1839) 442 526 

Sphaerozetes piriformis (Nicolet 1855) 164 305 306 319 567 



Svalbardia paludicola Thor 1930 181 318 442 563 567 

Trichoribates novus (Sellnick 1928) 56 567 

Trichoribates setiger (Trägårdh 1910) 235 442 567 

Trichoribates trimaculatus (C.L.Koch 1836) 56 182 219 555 567 

Cymbaeremaeoidea 

Achipterioidae 

Achipteria sp. 442 

Parachipteria punctata (Nicolet 1855) 441 442 526 567 

Micreremidae 

Micreremus brevipes (Michael 1888) 235 442 567 

Damaeoidea 

Damaeidae 



Subbelba montana (Kulcyznski 1992) 

Oribata ursina Thor 1930 

Spatiodamaeus sp. 

?Dameus sp 

219 318 319 567 

235 445 

Damaeus onustus C.L. Koch, 1841 526 567 

Euphthiracaroidea 

Euphthiracaridae 

Microtritia sp. 272 

Galumnoidea 

Galumnidae 

Pergalumna nervosa (Berlese 1914) 272 442 567 

Gustavioidea 

Peloppiidae 

Ceratoppia bipilis (Hermann 1804) 

Eremaeus bipilis (Hermann) 

442 

41 42 121 164 305 306 319 555 567 



Ceratoppia hoeli Thor 1930 13 25 66 69 70 152 181 182 235 377 411 420 421 

442 501 567 

Ceratoppia sphaerica (L.Koch 1879) 

Notaspis bipilis var. sphaerica (L. 

Koch) 

Gymnodamaeoidea 

Gymnodamaeidae 

Gymnodamaeus sp. 272 

25 219 306 335 442 526 563 567 

Hermannioidea 

Hermanniidae 

Hermannia reticulata Thorell 1872 13 34 66 69 152 164 181 182 219 235 305 318 319 

320 

567 

335 342 377 387 411 420 421 442 501 526 555 563 

Hermannia scabra (L.Koch 1879) 182 319 563 567 

Licneremaeoidea 

Licneremaeidae 

Scutovertex minutus (C.L.Koch 1835) 181 182 235 442 567 

Nanhermannioidea 


420


Nanhermannioidae 

Nanhermannia sellnicki (Forsslund 1958) 442 567 

Nothroidea 

Camisiidae 

Camisia anomia Colloff 1993 13 66 69 335 365 375 377 387 411 418 420 421 458 

563 567 

Camisia biverrucata (C.L.Koch 1839) 

Nothrus biverrucatus C.L.Koch 

181 182 320 442 567 

Camisia borealis (Thorell 1871) 70 150 181 182 235 319 442 526 567 

Camisia dictyna Colloff 1993 567 

Camisia foveolata Hammer 1955 56 121 182 271 280 563 567 

Camisia horrida (Hermann 1804) 

Nothrus horridus (Hermann) Nothrus borealis Thorell 

34 181 182 219 235 318 320 342 442 567 



Camisia invenusta (Michael 1888) 

Nothrus invenusta (Michael) 

Camisia lapponica (Trägårdh 1910) 56 442 567 

Camisia spinifer (C.L.Koch 1836) 

Nothrus spinifer C.L.Koch 

567 

181 182 320 442 567 

Nothridae 

Nothrus palustris (C.L.Koch 1839) 441 442 567 

Platynothrus peltifer (C.L.Koch 1839) 

? Platynothrus punctatus (C.L.Koch) 

181 182 235 442 567 

Platynothrus punctatus (C.L.Koch 1839) 441 442 526 563 567 

Platynothrus septentrionalis Sellnick 1944 

Platynothrus capillatus (Berlese) 

var. septentrionalis Sellnick 

Oppioidea 

Autognetidae 

181 182 442 



Autogneta kaisilai Karppinen 1997 181 182 342 442 567 

Oppiidae 

Berniniella sp. 457 

Lauroppia fallax (Paoli 1908) 

Oppiella fallax (Paoli) Oppia fallax (Paoli) 

Dissorhina ornata (Oudemans 1900) 

Cosmoppia ornata (Oudemans) 

Oppia ornata (Oudemans) 

Microppia minus (Paoli 1908) 

Oppiella minus 

Dameosoma ornatum (Oudemans) 

181 182 320 342 442 567 

181 182 219 318 320 442 563 567 

272 442 567 

Oppiella neerlandica (Oudemans 1902) 442 457 595 

Oppiella nova Oudemans 1902 

Dameosoma neerlandicum 

(Oudemans 1900) 

Oppia neerlandica (Oudemans) 

Oppia nova (Oudemans) 

5 69 157 181 182 219 272 318 320 335 342 377 387 

411 421 441 442 526 567 595 


595


Oppiella (Moritzoppia) unicarinata (Paoli 1908) 442 457 

Moritzoppia neerlandica (Oudemans 1900) 

Oppia translamellata (Willmann) 

Moritzoppia splendens (C.L. Koch 1840) 

Discoppia splendens (C.L. Koch 

1840) 

70 121 181 182 191 219 235 271 272 320 342 398 411 

441 442 526 563 567 

442 526 567 

Suctobelbella hammeri (Krivolutsky 1966) 441 442 567 

Suctobelbella sarekensis (Forsslund 1941) 

Suctobelba sarekensis (Forsslund) 

Suctobelbella subcornigera (Forsslund 1941) 

Suctobelba subcornigera (Forsslund) 

181 182 342 442 567 

181 182 342 441 442 526 567 

Oribatelloidea 

Oribatellidae 

Oribatella arctica Thor 1930 181 318 320 501 561 567 

Oripodoidea 

Oribatulidae 



Oribatula tibialis (Nicolet 1885) 56 219 442 555 563 567 595 

Oribatula venusta Berlese 1908 

Zygoribatula venusta (Berlese) 

Phauloppia lucorum (C.L.Koch 1841) 

Notaspis lucorum (C.L.Koch) 

Michael 1897 

Zygoribatula exilis (Nicolet 1855) 

Oribatula exilis (Nicolet) 

56 181 182 235 320 342 442 501 567 

181 442 567 

164 182 305 318 319 442 567 

Oripodiidae 

Liebstadia similis (Michael 1888) 219 441 442 567 

Trombidiformes (Actinedida) 

Bdelloidea 

Bdellidae 

Bdella longicornis (Linnaeus 1758) 

Bdella decipiens Thorell 

Bdella semiscutata Thor 1930 318 320 

159 219 305 306 318 319 320 



Biscirus lapidarius (Kramer 1881) 

Scirus lapidarius (Kramer) 

20 318 

Cyta latirostris (Hermann 1804) 219 318 320 

Cyta brevirostris (L.Koch 1879) [a variety of C. latirostris] 

Ammonia brevirostris L.Koch sensu 

Trägårdh 

Neomolgus capillatus (Kramer 1881) 

Molgus capillatus (Kramer) 

Neomolgus littoralis (Linnaeus 1758) 

Bdella arctica Thorell 1872 

Bdella groenlandicus Trägårdh 

Bdella sanguinea (Trouessart) 

Bdella littoralis (Linnaeus) 

Molgus littoralis (L.) 

Neomolgus pallipes (L.Koch) 159 305 306 

Cunaxidae 

Cunaxoides croceus (C.L.Koch 1838) 

Eupalus croceus (C.L.Koch) 

Eupodoidea 

Eupodidae 

318 

12 20 41 159 219 305 306 318 411 465 558 563 

320 



Cocceupodes clavifrons (R. Canestrini1886) 

Coccorhagidia clavifrons 

(Canestrini) 

Eupodes clavifrons Canestrini 

219 318 320 

Eupodes variegatus C.L.Koch 1838 219 318 320 

Protereunetes borneri Thor 1934 219 320 

Penthaleidae 

Penthaleus maior (Duges 1834) 

Penthaleus insulanus Thorell 

Penthalodidae 

Penthalodes ovalis (Dugès 1834) 

Chromotydeus arcticus Trägårdh 

1905 

Penthaleus arcticus Trägårdh 

Rhagidiidae 

Coccorhagidia clavifrons (Canestrini1886) 

Rhagidia clavifrons (Canestrini) 

Penthalodes ovalis Dugès 

219 318 319 320 

219 306 318 319 320 

Rhagidia gelida Thorell 1872 20 41 159 219 305 306 318 319 320 

Lebertioidea 

318 



Sperchonidae 

Sperchon brevirostris (Koenike 1895) 

Sperchon lineatus Thor 1899 

Pachygnathoidea 

Alicorhagiidae 

Alicorhagia clavipilus (Thor 1931) 

Epistomalycus clavipilus Thor 

Alicorhagia plumipilis (Thor 1931) 

Epistomalycus plumipilis Thor 

Nanorchestidae 

Nanorchestes arboriger (Berlese 1904) 

Alycus arboriger Berlese 

Pygmephoroidea 

Pygmephoridae 

Bakerdania arcticus (Thor 1934) 

Pediculoides arcticus (Thor) 

Raphignathoidea 

Stigmaeidae 

Eustigmaeus oudemansi (Thor 1930) 

Ledermülleria oudemansi (Thor) 

20 305 318 

320 

320 

219 318 320 

320 

318 



Eustigmaeus pulchellus (Thor 1930) 

Liostigmaeus pulchellus Thor 

Tetranychoidea 

Tetranychidae 

Bryobia borealis Oudemans 1930 306 318 320 

Trombidioidea 

Podothrombiidae 

Podothrombium bicolor (Hermann 1804) 318 

Podothrombium curtipalpe (Thor 1900) 318 

Podothrombium svalbardense Oudemans 1930 318 

Tydeoidea 

Tydeiidae 

Coccotydeus globifer (Thor 1931) 320 

Microtydeus constans Thor 1931 219 320 

318 



Triophtydeus pinicolus (Oudemans 1929) 320 

Tydaeolus tenuiclaviger (Thor 1931) 

Coccotydeus tenuiclaviger (Thor) 

Tydeus croceus (Linnaeus 1758) 

Tydeus langei Thor 1934 320 

Tydeus svalbardensis Thor 1932 219 320 

Parasitiformes 

Astigmata 

Analgoidea 

Avenzoariidae 

Zachvatkinia isolata Mironov, 1989 565 

Gamasida (Mesostigmata) 

Ascoidea 

Ascidae 

Antennoseius (Vitzthumia) oudemansi (Thor 1930) 

Vitzthumia oudemansi Thor 

320 

56 318 320 543 564 563 570 591 595 



Arctoseius babenkoi Makarova 1999 450 451 452 543 

Arctoseius cetratus (Sellnick 1940) 452 543 591 

Arctoseius haalovi Lindquist 1963 452 564 563 571 575 591 595 

Arctoseius laterincisus Thor 1930 318 320 543 563 

Arctoseius multidentatus Evans 1955 56 452 543 564 570 591 595 

Arctoseius ornatus Evans 1955 450 452 543 591 

Arctoseius pristinus Karg 1992 56 543 

Arctoseius tajmricus Petrova and Makarova 1991 56 450 452 462 543 570 591 

Arctoseius tschernovi Makarova 2000 564 591 



Arctoseius weberi Evans 1955 56 452 543 564 591 

Melichares sp. 56 543 

Halolaelaps (Halogamasellus) gerlachi (Hirschmann, 1966) 591 

Proctolaelaps parvanalis (Thor 1930) 

Lasioseuis parvanalis Thor 

Zerconopsis labradorensis Wvans et Till, 1960 591 

Zerconopsis sp. 56 543 

Phytoseiidae 

Amblyseius magnanalis Thor 1930 

Lasioseuis magnanalis Thor 

Dermanyssoidea 

Dermanyssidae 

Dermanyssus hirundinis (Hermann, 1804) 590 

Haemogamasidae 

318 320 542 543 564 591 

318 320 543 564 591 594 



Haemogamasus ambulans (Thorell 1872) 

Eulaelaps ambulans (Thorell) 

201 318 320 564 591 

Hypoaspididae 

Hypoaspis ovalis L.Koch 159 305 319 543 

Laelapidae 

Laelaps hilaris (Koch) 201 543 591 

Macrochelidae 

Macrocheles muscaedomesticae (Scopoli, 17772) 591 

Eviphidoidea 

Eviphididae 

Thinoseius spinosus (Willmann, 1939) 525 563 573 591 

Parasitoidea 

Parasitidae 

Pergamasus sp. 219 

Paragamasus (Aclerogamasus) insertus (Micherdziński, 1969) 580 595 



Vulgarogamasus immanis (Berlese, 1904) 579 

Vulgarogamasus remberti (Oudemans 1912) 580 595 

Rhodacaroidea 

Ologamasidae 

Gamasellus borealis (Koch 1879) 306 543 

Veigaioidea 

Veigaiidae 

Zercon andrei Sellnick, 1958 574 591 

Zerconoidea 

Zerconidae 

Zercon forsslundi Sellnick 1958 56 543 564 570 591 

Zercon solenites Haarløv 1942 564 563 572 570 591 

Zercon spinosus 



Zercon triangularis C.L.Koch 1836 318 320 543 591 

Zercon zelawaensis Sellnick 1944 56 543 591 

Zerconopsis muestairi (Schweizer, 1949) 574 

Ixodida (Metastigmata) 

Ixodoidea 

Ixodidae 

Ixodes (Ceratixodes) uriae White 1852 

Araneae 

_ 

_ 

_ 

Ixodes putus (Pickard-Cambridge) 

Gnaphosidae 

Micaria constricta Emerton 1894 

Micaria eltoni Jackson 

41 554 596 

Hahniidae 

Hahnia helveola Simon 1875 138 169 312 

Linyphiidae 

138 146 160 166 167 169 306 314 



Halorates holmgreni (Thorell 1871) 

Coryphaeolanus holmgreni Thor 

Coryphaeus mendicus L. Koch 

Collinsia holmgreni (Thorell 1871) 

Halorates spetsbergensis (Thorell 1871) 

Collinsia spetsbergensis (Thorell 

1871) 

? Typhochraestus borealis Thor 

Microeriogone spetsbergensis 

(Thorell) 

Halorates thulensis (Jackson 1934) 

Collinsia thulensis (Jackson 1934) 

Lophomma holmgreni (Thorell) 

Coryphaeus holmgreni Thor 

Typhochraestus spetsbergensis 

(Thor) 

Erigone spetsbergensis Thorell 

Erigone longipalpis Sundevall 1830 319 

Erigone arctica palearctica Braendegaard 1934 

Erigone arctica White 

41 42 43 44 120 137 146 160 163 166 167 168 169 

305 306 312 322 570 

43 56 120 137 138 146 160 161 129 163 166 167 168 

169 199 271 305 306 312 314 318 319 345 378 420 421 

535 570 

146 160 312 

12 43 55 56 120 137 138 146 160 129 163 166 167 

168 169 199 271 305 306 312 314 318 345 378 411 420 

421 535 570 595 

Erigone psychrophila (Thorell 1871) 137 129 138 146 160 163 166 167 168 199 306 312 345 

378 420 421 535 570 


314 

312


Erigone tirolensis L. Koch 1872 20 41 42 43 44 160 166 167 169 305 306 312 322 

535 570 

Hilaria glacialis (Thorell 1871) 

Erigone glacialis Thorell 

Improphantes complicatus (Emerton 1882) 

Lepthyphantes hyperboreus Strand 

Lepthyphantes complicatus 

(Emerton 1882) 

Mughiphantes sobrius (Thorell 1871) 

Micryphantes hyperboreus C.L.Koch 

Linyphia sobria Thorell Lepthyphantes sobrius (Thorell 1871) 

Mecynargus borealis (Jackson 1930) 

Rhaebothorax borealis (Jackson) Conigerella borealis (Jackson) 

Agyneta nigripes (Simon 1884) 

Agyneta nigripes (Simon) 

Meioneta nigripes (Simon 1884) 

Micryphantes nigripes Simon 

43 55 56 120 137 138 146 160 129 163 166 167 168 

169 

570 

199 305 306 312 314 318 319 345 378 420 421 535 

43 120 160 169 312 345 

43 120 129 137 138 146 160 163 166 167 168 169 199 

305 306 312 314 319 345 535 570 

138 146 160 191 193 194 199 

41 42 44 146 160 161 163 167 169 199 306 312 314 

322 345 378 420 421 535 570 


318 

535 

570


Oreonetides vaginatus (Thorell 1872) 312 

Walckenaeria karpinskii (O.P.-Cambridge 1873) 

Cornicularia karpinskii (Cambridge) 

Tapinocyba insecta (L.Koch 1869) 

Colobocyba insecta L.Koch 

Philodromidae 

Thanatus formicinus (Clerck 1757) 345 

146 160 322 

169 312 


Mandibulata 

Collembola 

Arthropleona 

_ 

_ 


Arrhopalitidae 

Arrhopalites principalis Stach 1945 

Arrhopalites binoculatus (Borner) 

Entomobryidae 

Entomobrya nivalis (L. 1758) 

Entomobrya subarctica Stach 1962 

Lepidocyrtus lignorum Fabricus 1793 

Lepidocyrtus lanuginosus (Gmelin) 

Hypogastruridae 

Hypogastrura assimilis Krausbauer, 1898 580 595 

Bonetogastrura nivalis (Martynova 1973) 117 

56 112 114 116 318 326 327 387 562 

33 112 116 286 326 327 562 

58 60 69 116 112 117 211 284 286 306 318 319 326 

342 387 562 587 



Ceratophysella longispina (Tullberg 1876) 

Hypogastrura longispina (Tullberg) 

Hypogastrura armata (Nicolet) 

Ceratophysella succinea (Gisin 1949) 

Hypogastrura hirsuta Valpas 

Achorutes longispinus 

Hypogastrura succinea Gisen Hypogastrura armata (Nicolet) 

Hypogastrura concolor (Carpenter 1900) 112 116 117 563 570 

Hypogastrura purpurescens (Lubbock 1868) 580 595 

Hypogastrura sensilis (Folsom 1919) 112 116 570 

Hypogastrura tullbergi (Schäffer 1900) 

Achorutes dubius Tullberg 

Hypogastrura spitsbergensis Stach 

Achorutes tullergi Schäffer 

58 60 107 112 116 117 140 141 142 143 211 263 266 

278 286 306 318 319 326 327 358 400 418 420 421 443 

444 470 537 562 

327 

33 112 114 116 122 123 219 284 326 327 562 

13 29 31 32 33 56 58 66 68 69 70 98 112 

116 117 123 150 152 155 156 157 271 211 241 262 263 

264 266 268 286 296 306 284 326 327 352 354 358 363 

373 375 375 387 397 418 420 421 444 445 537 562 563 

570 156 



Hypogastrura viatica (Tullberg 1871) 

Achorutes viaticus Tullberg 

Podura hyperborea Boheman 1865 

Hypogastrura or Ceratophysella. 

Species unknown. 

20 56 58 60 109 112 114 116 117 123 140 141 142 

143 191 193 194 211 212 256 263 266 278 284 286 305 

306 308 318 319 326 327 332 353 358 411 418 420 421 

429 434 434 444 454 470 472 492 537 539 558 562 563 

570 587 595 256 

112 116 327 562 

Willemia anophthalma Börner 1901 112 114 116 122 219 263 266 318 326 327 562 570 

Willemia scandinavica Stach 1949 112 114 116 117 141 143 266 284 562 563 570 

Willemia similis Mills 1934 114 116 117 562 570 

Xenylla humicola (Fabicius 1780) 20 41 42 56 58 60 110 112 114 116 117 122 141 

143 211 263 266 278 284 286 305 306 318 319 326 327 

332 358 470 537 562 570 

Isotomidae 


286


Agrenia bidenticulata (Tullberg 1876) 

Isotoma bidenticulata Tullberg 

Archisotoma besselsi (Packard 1877) 

?Isotoma spitzbergensis Lubbock 

Isotoma janmayensis (Wahlgren) 

?Isotoma arctica Stscherbakow 

15 20 56 58 60 112 114 116 117 208 211 212 263 

266 271 278 286 306 318 319 326 327 537 562 570 

58 60 112 114 116 117 122 211 217 219 266 278 305 

306 327 562 

Archisotoma megalops (Bagnall 1939) 112 114 116 127 208 209 562 

Archisotoma polaris Fjellberg and Poinsot 1975 

Isotoma arctica Stscherbakow 

Folsomia coeruleogrisea (Hammer, 1938) 

Isotomina gracilis Stach 

Folsomia alpha Grow and 

Christiansen 1976 

Folsomia bisetosa Gisin 1953 

Folsomia fimetaria (L.) 

Folsomia sensibilis Kseneman 

Folsomia gracilis (Stach 1962) 

Folsomia bisetosella Fjellberg 2005 523 562 563 

13 116 117 118 562 570 

33 42 69 70 112 116 117 122 141 143 219 249 286 

326 327 537 112 114 562 563 570 

41 58 60 69 112 114 116 117 122 123 141 143 219 

249 263 266 271 284 306 318 319 326 327 358 377 387 

420 421 470 562 563 570 595 


319 

563 

318


Folsomia ciliata Babenko & Bulavintsev, 1993 562 

Folsomia quadrioculata (Tullberg 1871) 

Isotoma quadrioculata Tullberg 

Folsomia binoculata (Wahlgren, 1899) 

Isotoma binoculata Wahlgren Folsomia regularis Hammer 1953 

Folsomia fimetaria (Linnaeus, 1758) 580 595 

16 20 29 32 37 56 58 60 68 69 109 111 112 

114 116 117 123 140 141 142 143 191 193 194 211 212 

217 243 249 262 266 268 271 278 284 305 306 318 319 

326 327 327 358 363 373 375 377 387 400 411 418 420 

421 

595 

428 429 443 444 470 501 142 558 562 563 570 587 

33 58 60 112 116 117 249 271 284 286 318 319 326 

327 562 570 

Folsomia sexoculata (Tullberg 1871) 20 58 60 114 116 122 140 141 142 143 219 249 286 

305 306 318 326 327 400 429 443 444 539 562 570 

Folsomia stella Grow and Christiansen 1976 69 114 116 562 595 

Folsomia taimyrica Martnova 1973 56 114 116 326 562 570 


429


Parisotoma notabilis Schäffer 1896 

Isotoma notabilis Schäffer 1896 

Isotoma anglicana Lubbock 1862 

Isotoma viridis Bourlet Isotoma palustris (Gmelin) 

Desoria grisea (Lubbock, 1869) 580 595 

Desoria multisetis (Carpenter and Phillips 1922) 

Isotoma multisetis Carpenter and 

Phillips 1922 

Desoria neglecta (Schäffer 1900) 

Isotoma fennica (Reuter) 

Isotoma neglecta Schäffer 1900 

Desoria olivacea (Tullberg 1871) 

Isotoma olivacea Tullberg 1871 

Isotoma regularis Hammer 1957 

Isotoma binoculata Wahlgren 

Isotoma violacea Tullberg 

112 114 116 122 219 284 562 595 

15 20 41 42 56 58 60 69 70 112 114 116 117 

122 211 212 219 263 266 278 284 286 305 306 318 319 

326 327 332 358 377 387 411 418 420 421 470 502 537 

558 562 562 595 

278 

20 58 60 112 116 305 327 562 

20 

562 

33 58 60 112 114 116 117 263 286 318 319 327 

58 112 114 116 263 266 306 318 562 

112 562 



Desoria fennica (Reuter, 1895) 

Isotoma ruseki Fjellberg 1979 

Desoria tshernovi Martynova 1974 

Isotoma fennica (Reuter) 

Isotoma olivacea (Tullberg) 

Isotoma nanseni Fjellberg 

Isotoma tshernovi 

Pseudanurophorus alticola Bagnall, 1949 595 

Pseudanurophorus inoculatus Bödvarsson 1958 562 

112 116 562 

56 69 112 116 117 326 327 418 537 562 595 

Pseudanurophorus psammophilus Potapov & Stebaeva, 2002 56 112 114 116 117 122 326 562 

Tetracanthella arctica Cassagnau 1959 

Tetracanthella wahlgreni 

Linnaniemi: Thor 

Tetracanthella coerulea (Haller) 

Tetracanthella pilosa Schött: 

Wahlgren 

Vertagopus arcticus Martynova 1969 112 117 562 

Vertagopus pseudocinereus Fjellberg 1975 112 116 562 

20 58 60 112 114 116 123 141 143 271 305 318 319 

327 332 358 375 387 418 458 470 562 587 


458


Neanuridae 

Anurida granaria (Nicolet 1847) 58 112 116 306 318 327 562 

Anurida maritima (Guerin 1836) 117 562 

Anurida polaris (Hammer 1954) 

Anurida fridgida Fjellberg 

Friesea quinquespinosa Wahlgren 1900 

Friesea nauroisi Cassagnau 

Micranurida pygmaea Börner 1901 

Anurida pygmaea (Börner) 

Odontellidae 

Xenyllodes armatus Axelson 1903 112 116 318 327 562 

Onychuiridae 

Deuteraphorura variablis (Stach 1954) 580 595 

56 69 70 112 114 116 117 141 143 263 266 284 326 

537 562 570 

112 116 117 284 326 327 358 470 562 563 570 595 

41 56 112 114 116 117 122 219 263 284 326 387 537 

562 570 



Megaphorura arctica (Tullberg 1876) 

12 13 20 35 41 42 57 58 60 66 69 109 112 

Aphorura arctica Tullberg 

Lipura arctica Tullberg 

114 116 117 122 150 151 152 153 155 156 157 162 194 

Onychuirus armatus (Tullberg) var. Onychiurus arcticus (Tullberg) 

211 217 219 243 263 266 281 282 284 286 296 305 306 

arcticus Tullberg 

Protaphorura arctica 

307 

375 

318 

398 

318 

408 

326 

409 

327 

411 

332 

418 

334 

425 

340 

152 

342 

470 

344 

473 

355 

477 

358 

498 

363 

501 

503 532 539 552 558 562 563 570 577 584 585 587 589 

594 597 598 600 601 602 603 604 605 

Mesaphorura jirii Rusek 1982 112 114 116 562 

Mesaphorura macrochaeta Rusek 1976 

Tullbergia krausbaueri Börner: Gisin 

112 114 116 117 122 219 562 

Mesaphorura tenuisensillata Rusek 1974 112 114 116 562 

Oligaphorura groenlandica (Tullberg 1876) 

Onychiurus groenlandicus (Tullberg) Lipura groenlandica (Tullberg) 

Oligaphorura ursi (Fjellberg 1984) 

Onychiurus ursi (Fjellberg) 

13 56 58 60 66 69 112 114 115 116 117 122 123 

140 141 143 152 156 157 193 211 217 219 263 266 281 

282 284 306 318 319 326 327 358 373 375 377 400 411 

418 420 420 429 443 444 470 537 539 558 562 563 570 

595 193 

114 115 116 117 387 562 563 570 595 



Thalassaphorura duplopunctatus (Strenzke 1954) 

Onychiurus duplopunctatus 

(Strenzke) 

Onychuirus neglectus Schäffer 

Protaphorura duplopunctatus 

(Strenzke 1954) 

Protaphorura macfadyeni (Gisin 1953) 

Aphorura armata Tullberg: 

Wahlgren 

Onychiurus duodecimpunctatus 

Onychiurus debilis Moniez: (Gisin) 

Aphorura neglecta Schäffer: 

Wahlgren 

Onychuirus macfadyyeni (Gisin) 

Protaphorura subuliginata Gisin 1956 562 563 595 

60 112 114 116 117 122 219 284 319 326 327 332 562 

41 42 112 114 116 122 219 326 332 562 

Tullbergia arctica Wahlgren 1900 112 114 116 122 219 326 327 562 

Chaetophorura simplex (Gisin 1958) 

Neelipleona 

_ 

_ 

Tullbergia simplex Gisin 1958 

Neelidae 

Megalothorax minimus (Willem 1900) 

Neelus (Megalothorax) minimus 

Willem: Gisin 

112 114 116 117 562 

112 114 116 117 122 219 263 326 562 



Symphypleona 

_ 

_ 

Katiannidae 

Sminthurinus aureus (Lubbock 1862) 20 112 318 327 562 

Sminthurinus concolor (Meinert 1896) 

Sminthurinus niger (Lubbock) 

Sminthurididae 

Sminthurides inaequalis armatus Bretfeld 2000 359 420 421 562 

Sminthurides malmgreni (Tullberg 1876) 

Sminthurus malmgreni Tullberg 

Sphaeridia pumilis (Krausbauer 1898) 112 116 562 

20 33 41 42 58 60 69 112 114 116 117 122 211 

219 263 286 305 306 284 318 326 327 377 387 420 421 

562 570 595 

15 20 56 58 60 112 116 117 200 266 278 286 305 

318 319 327 358 418 470 562 


318 

562

Insecta 

Endopterygota 

Siphonaptera 

_ 

_ 


Ceratophyllidae 

Ceratophyllus vagabundus vagabundus (Boheman 1866) 

Pulex vagabunda Boheman Ceratophyllus digitalis (Wahlgren) 

Mioctenopsylla arctica arctica Rothschild 1922 

Coleoptera 

_ 

_ 

Mioctenopsylla arctica Rothschild 

Anthicidae 

Anthicus flavipes (Panzer 1797) 179 

75 105 176 223 319 563 

71 128 176 223 555 563 

Byrrhidae 

Simplocaria metallica (Sturm 1807) 113 128 261 293 303 305 

Carabidae 

Amara quenseli (Schönherr 1806) 17 113 302 303 305 411 



Chrysomelidae 

? Gonioctena arctica Mannerheim 1853 104 

Phytodecta ? 104 306 

Coccinellidae 

Coccinella septempunctata L. 1758 

Cryptophagidae 

Atomaria atricapilla Stephens 1830 113 179 

Atomaria lewisi Reitter 1877 113 178 

Curculionidae 

Orchestris saliceti Fabricus 

Tachyerges (Orchestris) saliceti 

Fabricus 

Isochnus flagellum (Erichson 1902) 

? Rhynchaenus foliorum (Müller) Rhynchaenus flagellum (Ericson 

1902) 

Dermestidae 

76 104 108 198 306 319 

113 128 146 178 303 305 318 387 



Reesa vespulae (Milliron, 1939) 

Dytiscidae 

Hydroporus striola (Gyllenhal 1827) 196 

Lathridiidae 

Latrididius minutus (L. 1767) 

Enicmus minutus L. 

Silvanidae 

Oryzaephilus mercator (Fauvel 1889) 370 

Staphylinidae 

Atheta graminicola (Gravenhorst 1806) 

? Atheta prope frigida Sahlberg 1806 

Boreophilia subplana (J. Sahlberg 1880) 

Atheta subplana J. Sahlberg 

Eucnecosum brachypterum (Gravenhorst 1802) 

Arpedium brachypterum 

(Gravenhorst) 

20 105 205 

12 56 76 104 128 146 156 157 408 411 501 558 595 

76 113 128 146 198 319 

20 113 198 205 


Diptera 


Micralymma marinum (Strøm 1783) 20 76 113 146 198 205 319 

Olophrum boreale (Paykull 1792) 20 41 113 205 

Omalium caesum Gravenhorst 1806 113 

Omalium septentrioneis Thomson 1857 20 205 

Brachycera 

_ 

Anthomyiidae 

Fucellia focorum (Fallén 1819) 

Scatomyza hyperborea Boheman 

Zaphne frontata (Zetterstedt 1838) 

Acroptena frontata Zetterstedt 

Scatomyza obscura Boheman 

Calliphoridae 

Calliphora vicina Robineau-Desvoidy 1830 

Calliphora erythrocephala (Meigen 

1826) 

Aricia frontata Zetterstedt 

Hylemyia frontata Zetterstedt 

146 321 

42 65 120 305 306 308 319 321 

306 



Cynomyia mortuorum (L. 1761) 20 41 42 101 219 553 

Protophormia atriceps (Zetterstedt, 1845) 

Boreellus atriceps (Zetterstedt, 1845) 

Protophormia terranovae (Robineau-Desvoidy 1830) 

Phormia terrae-novae Robineau- 

Desvoidy 1830 

Phormia groenlandica (Zetterstedt) 

Coelopidae 

Coelopa eximia Stenhammer 1854 319 

Fucomyia frigida Fallén 1805 

Coelopa frigida (Fabricus) 

306 529 

Drosophilidae 

Drosophila funebris (Fabricus 1787) 125 281 388 

105 146 158 239 306 423 505 

59 65 105 125 146 306 319 505 

Empididae 

Rhamphomyia caudata Zetterstedt 1838 65 125 146 185 187 305 306 319 505 

Heleomyzidae 



Heleomyza modesta (Meigen, 1878) 156 232 319 341 

Heleomyza serrata (L. 1758) 20 101 105 185 187 

Heleomyza borealis (Boheman 1865) 

Leria modesta Meigen (det. Collin) ? Leria geniculata Zetterstedt 

Helomyzidae 

Leria geniculata Zetterstedt 1847 120 319 

Neolaria prominens (Becker 1897) 125 267 505 

Neoleria septentrionalis (Collin 1923) 

Leria septentrionalis Collin (det. 

Collin) 

Muscidae 

Spilogona brunneifrons Ringdahl 1931 313 321 

Spilogona denudata (Holmgren 1869) 

Aricia denudata Holmgren Aricia ranunculi Holmgren 

125 497 498 505 541 552 600 

20 59 65 101 319 

319 321 



Spilogona dorsata (Zetterstedt 1845) 

Aricia conspurcata Holmgren 

Aricia fuliginosa Holmgren 

Aricia illota Holmgren 

Limnophora hyperborea Boheman 

Spilogona megastoma (Boheman 1866) 

Aricia ludibunda Holmgren 

Limnophora arctica Becher 

Aricia dorsata Zetterstedt 

Aricia hyperborea Boheman 

Aricia labiosa Boheman 

Aricia pauxilla Holmgren 

Limnophora megastoma (Boheman) 

Spilogona quinquelineata (Zetterstedt 1838) 313 321 

Spilogona triangulifera (Zetterstedt 1838) 

Aricia triangulifera Zetterstedt 

Limnophora triangulifera Zetterstedt 

Aricia vitticollis Zetterstedt 

Spilogona varsviensis Schnabl in Schnabl and Dziedzicki 1911 

Piophilidae 

Parapiophila vulgaris (Fallén 1820) 

Piophila vulgaris Fallén Piophila sp. cf. vulgaris Fallén 

Scatophagidae 

Allomyella frigida (Holmgren 1883) 

Microprosopa frigida (Holmgren) 

20 65 101 102 219 305 306 319 505 

59 65 305 319 321 505 

59 120 185 187 319 321 

20 101 213 505 

20 101 



Scatophaga furcata (Say 1823) 

Scatophaga squalida Meigen 

Scatophaga litorea (Fallén 1819) 

Scatophaga nigripes (Holmgren) 

59 125 146 319 411 420 

20 59 101 125 146 219 

Scatophaga varipes (Holmgren 1883) 65 219 305 306 

Sphaeroceidae 

Copromyza (Borborus) uncinatus (Duda 1923) 125 313 

Norrbomia fumipennis (Stenhammar 1855) 

Copromyza (Borborus) fumipennis 

Stenhammer 

125 319 505 

Syrphidae 

Syrphus ribesii (L. 1758) 65 105 120 146 156 174 177 185 

Syrphus torvus Osten-Sacken 1875 105 106 281 282 



Parasyrphus tarsatus (Zetterstedt 1838) 

Scaeva dryadis Holmgren 

Syrphus tarsatus Zetterstedt 

Syrphus tarsatus Zetterstedt 

(dryadis Holmgren) 

Tachinidae 

Tachina glacialis Boheman 319 

Nematocera 

_ 

Culicidae 

Aedes (Ochlerotatus) nigripes (Zetterstedt 1838) 

Aedes alpinus L. Culex nigripes Zetterstedt 

Mycetophilidae 

Allodida sp. 146 

Boletina maculata Holmgren 1870 

Boletina apicalis (Walker) 

Coelosia tenella (Zetterstedt 1852) 

Boletina setipennis (Holmgren) 

265 420 421 

59 96 146 147 282 296 305 319 422 

20 59 96 98 101 146 319 323 

59 96 98 146 319 323 



Exechia frigida (Boheman 1865) 

Mycetophila frigida Boheman 

Phronia egregia Dziedzicki1889 

Phronia melaena Edwards 

Rhymosia sp. 146 

Scatopsidae 

Scatopse notata (L. 1758) 

Tipula notata L 

Sciaridae 

Bradysia nervosa (Meigen 1818) 

Sciara variabilis Zetterstedt 

Bradysia praecox (Meigen 1818) 

Sciara praecox Meigen 

Camptochaeta consimilis (Holmgren 1869) 

Corynoptera consimilis (Holmgren) Sciara consimilis Holmgren 

20 59 96 97 101 121 219 305 

98 146 306 323 

99 

308 

96 98 305 306 

20 96 101 323 



Camptocheata delicata (Frey 1948) 

Neosciara delicata Lengersdorf Sciara pallidiventris Holmgren 

Corynoptera röderi Lengersdorf 1931 

Sciara röderi Lengersdorf 

Lycoriella (Hemineurina) abbrevinervis (Holmgren 1869) 

Sciara abbrevinervis Holmgren 

Lycoriella (Hemineurina) cochleata (Rübsaamen 1898) 323 

Lycoriella (Hemineurina) modesta (Staeger 1840) 

Lycoriella arctica (Holmgren) Frey 

Sciara holmgreni Rübsaamen 

(frigida Holmgren) 

Sciara arctica Holmgren 

Sciara frigida Holmgren 

Sciara groenlandica Rübsaamen 

Sciara ecalcarata Holmgren 

Lycoriella postconspicua Mohrig, 1985 505 

Lycoriella (Hemineurina) vitticollis (Holmgren 1883) 

Sciara glacialis Rübsaamen Sciara permutata (Lundbeck) 

Lycoriella (Lycoriella) parva (Holmgren 1869) 

Sciara parva Holmgren 

20 96 98 99 101 305 306 319 505 

20 101 

96 323 

20 96 97 99 101 306 318 323 

318 323 



Lycoriella sp. 595 

Schwenckfeldina tridentata (Rübsaamen 1898) 

Rhynchosciara laguncularis 

Lengersdorf 

Simulidae 

Prosimulium ursinum (Edwards 1935) 

Simulium ursinum Edwards 

Sciara tridentata Rübsaamen 

20 96 97 101 102 305 306 318 505 

20 101 100 

Trichoceridae 

Trichocera borealis Lackschewitz 1934 73 74 296 310 

Trichocera hiemalis (De Geer 1760) 

incisurata 

Trichocera lutea (Becher 1886) 

Metatrichocera lutea (Becher) 

319 

20 73 74 96 97 98 101 110 219 

Trichocera maculipennis Meigen 1818 20 97 101 110 219 

Trichocera parva Meigen 1804 319 



Trichocera saltator (Harris 1776) 96 98 102 

Chironomidae 

Chironominae 

Chironomus (Chironomus) hyperboreus Staeger 1841 210 

Chironomus (Chironomus) lugubris Zetterstedt 1850 154 210 513 551 

Sergentia coracina (Zetterstedt 1850) 210 

Micropsectra insignilobus Kieffer 1924 557 

Micropsectra logani Johannsen 1928 557 

Micropsectra radialis Goetghebuer 1939 210 557 

Paratanytarsus austriacus (Kieffer 1924) 210 513 557 

Tanytarsus heliomesonyctios Langton 1999 557 



Diamesiinae 

Diamesa aberrata Lundbeck 1889 154 210 513 551 

Diamesa arctica (Boheman 1865) 154 210 513 563 

Diamesa arctica sensu Willassen and Velle (unpubl.) 551 

Diamesa bertrami Edwards 1935 154 210 513 551 

Diamesa bohemani Goetghebuer 1932 210 513 551 

Diamesa hyperborea Holmgren 1869 210 

Diamesa incallida (Walker) 513 

Diamesa lindrothi Goetghebuer 1931 154 210 

Diamesa lundstroemi Kieffer 1911 210 

Orthocladiinae 



Camptocladius stercorarius (De Geer 1776) 210 

Chaetocladius dissipatus (Edwards), 513 551 

Chaetocladius dentiforceps (Edwards) 513 551 

Chaetocladius holmgreni (Jabcobson 1898) 210 513 551 

Chaetocladius laminatus Brundin 1947 154 551 

Chaetocladius perennis (Meigen 1830) 210 513 551 

Cricotopus (Cricotopus) ephippium (Zetterstedt 1838) 210 

Cricotopus (Cricotopus) lestralis (Edwards 1924) 210 513 

Cricotopus (Cricotopus) pilosellus Brund 1956 210 



Cricotopus (Cricotopus) polaris Keiffer 1926 513 551 

Cricotopus (Cricotopus) tibialis (Meigen 1804) 210 

Cricotopus (Cricotopus) villosus Hirvenoja 1973 210 

Cricotopus (Isocladius) glacialis Edwards 1922 210 

Heterotrissocladius subpilosus (Kieffer 1911) 210 

Hydrobaenus conformis (Holmgren 1869) 210 513 551 563 

Limnophyes brachytomus (Kieffer 1922) 154 210 513 551 

Limnophyes edwardsi Sæther 1990 210 513 551 

Limnophyes eltoni (Edwards 1921) 210 513 



Limnopheys lapponicus Goetghebuer 1939 210 

Limnophyes pumilio (Holmgren 1869) 210 551 

Metriocnemus cataractarum Kieffer 1919 210 

Metriocnemus eurynotus (Holmgren 1883) 210 513 

Metriocnemus obscuripes (Holmgren 1869) 154 

Metriocnemus picipes (Meigen 1818) 210 

Metriocnemus similis Kieffer 1922 210 

Metriocnemus ursinus (Holmgren 1869) 210 513 551 

Oliveridia tricornis (Oliver 1976) 491 



Orthocladius (Eudactylocladius) gelidus Kieffer 1922 210 513 

Orthocladius (Euorthocladius) telochaetus Langton 1985 154 210 513 551 

Orthocladius (Euorthocladius) thienemanni Keiffer 513 551 

Orthocladius (Orthocladius) decoratus (Holmgren 1869) 210 513 551 

Orthocladius (Orthocladius) knuthi Soponis 1977 210 

Orthocladius (Orthocladius) obumbratus Johannsen 1905 210 

Orthocladius (Orthocladius) rhyacobius Kieffer 1911 513 

Orthocladius (Orthocladius) trigonolabis Edwards 1924 

Orthocladius (Pogonocladius) consobrinus (Holmgren 1869) 513 551 



Parachlus kiefferi 563 

Paraphaenocladius brevinervis (Holmgren 1869) 210 513 

Psectrocladius (Monopsectrocladius) calcaratus (Edwards 1929) 

Psectrocladius (Psectrocladius) barbimanus Edwards 1929 

Psectrocladius (Psectrocladius) borealis Kieffer 1919 210 

Psectrocladius (Psectrocladius) limbatellus (Holmgren 1869) 513 551 

Psectrocladius (Psectrocladius) octomaculatus Wülker 1956 

Psectrocladius (Psectrocladius) oxyura Langton 1985 21 

Psectrocladius (Psectrocladius) ventricosus Kieffer 1925 21 



Psectrocladius psilopterus (Keiffer 1906) 513 

Pseudosmittia oxoniana (Edwards 1922) 210 

Smittia brevipennis (Boheman 1856) 210 595 

Smittia extrema (Holmgren 1869) 210 

Smittia flexinervis (Kieffer 1911) 210 

Smittia lasiops (Malloch 1915) 210 

Zalutschia tatrica (Pagast 1935) 210 

Chironomus anthracinus - type 362 

Tanypodinae 

Arctopelopia barbitarsis (Zetterstedt 1850) 210 



Arctopelopia melanosoma (Goetghebuer 1933) 210 

Procladius crassinervis (Zetterstedt 1838) 

Hymenoptera 

Apocrita 

_ 

Braconidae 

Aphidius leclanti 578 

Biosteres sp. 20 255 

Diaeretellus svalbardicum Chaubert and Tomanović 2012 578 

Diaertiella rapae (M'Intosh 1855) 420 421 

Dendrocerus bifoveatus (Keiffer 1907) 378 387 417 420 421 

Ichneutes hyperboreus Holmgren 1869 305 306 319 333 378 420 421 



Trioxys sp. 378 

Ceraphronidae 

Conostigmus spetsbergensis (Holmgren 1869) 

Ceraphron spetsbergensis Holmgren 

Lagynodes pallidus Boheman 1832 331 

Cynipidae 

Alloxysta sp. 156 378 387 420 421 

Ichneumonidae 

Aclastus borealis (Boheman 1866) 

Hemiteles septentrionalis Holmgren 

Aclastus gracilis (Thomson 1884) 

? Aclastus gracilis minutus 

Bridgman 1886 

Aptesis erratica (Holmgren 1969) 

Microcryptus erraticus Holmgren Phygadeuon erraticus Holmgren 

319 

128 146 319 331 378 387 420 421 

20 255 

20 255 319 



Atractodes bicolor Gravenhorst 1829 var. arcticus Holmgren 

1872 

Atractodes pusillus (Förester 1876) 374 378 387 420 421 

Gelis glacialis (Holmgren 1869) 

Hemiteles glacialis Holmgren 

Hypamblys leucopgus (Holmgren 1869) 

Mesoleius leucopgus Holmgren 

Mesochorus palanderi Holmgren 1869 308 

Orthocentrus nigricornis Boheman 1866 

Stenomacrus nigricornis (Boheman) 

146 319 378 420 421 

319 

59 128 318 319 331 

Stenomacrus groenlandicus Jussila 1996 374 378 387 420 421 

Stenomacrus nigricornis (Boheman) 

Phygadeuon sp. 



Plectiscidea hyperborea (Holmgren 1869) 

Plectiscus hyperboreus Holmgren 

Stenomacrus pedestris (Holmgren 1869) 

Orthocentrus pedestris Holmgren 

Stenomacrus validicornis (Boheman 1866) 

Orthocentrus validicornis Boheman 

146 319 331 378 387 420 421 

Synodites lineiger (Thompson 1893) 378 380 420 421 

Tymmophorus obscuripes (Holmgren 1858) 

Diplazon arcticus (Holmgren) 

Symphyta 

_ 

Tenthredinidae 

Amauronematus hyperboreus (Thomson 1875) 259 319 

Amauronematus alberich Benson 1934 18 20 

20 128 146 305 318 319 331 333 

331 



Amauronematus villosus (Thomson 1862) 

Nematus villosus Thomson 

Amauronematus rubiginosus Hellén 1951 447 

Euura sp.? 318 

18 20 231 305 319 

Pontopristia amentorum (Förster 1854) 331 378 420 421 

Pristiphora frigida (Boheman 1865) 

Pristiphora adelungi Kenow Nematus frigidus Boheman 

Pristiphora stavdingeri (Ruthe 1859) 

Lepidoptera 

_ 

_ 

Pristiphora asperlatus Benson 

Noctuidae 

Apamea exulis (Lefèbvre, 1836) 

Apamea exulis (ef.) Apamea maillardi (Geyer 1834) 

18 20 59 128 146 259 305 306 

59 

4 175 204 455 



Syngrapha interrogationsis (L. 1758) 

Plusia interrogationsis (L.) 

Nymphalidae 

Nymphalis antiopa (L. 1758) 282 

204 269 

Vanessa cardui (L. 1758) 204 214 281 282 

Oecophoridae 

Hofmannospila pseudospretella (Stainton 1849) 

Papilionidae 

Parnassius apollo (L.1758) 72 

Pieridae 

Pieris napi (L.1758) 

Artogeia napi (L.) 

Margaritia sticticalis (L 1761) 

Pyralidae 

175 204 281 



Pempelia dilutella Hubner 1796 

Pempelia subornatella Duponchel 

Plutella xylostella (L. 1758) 

Plutella cruciferarum var. nivella ? Plutella maculipennis (Curtis) 

175 204 

Plutella polaris Zeller 1880 175 204 214 566 568 

Matilella fusca (Haworth 1811) 

Pyla fusca Haworth 1811 

Ephestia kuehniella Zeller 1879 

Trichoptera 

_ 

_ 

Limnophilidae 

Apatania zonella Zetterstedt 1840 

Exopterygota 

Anoplura 

_ 

_ 

Apatania arctica (Boheman) 

Goniataulius arcticus Boheman 

Apatelia zonella Zetterstedt 

20 41 104 110 175 204 205 214 253 281 282 287 306 

319 376 411 420 568 

204 382 379 

20 82 173 195 196 205 279 281 305 306 319 324 325 

378 491 538 



Echinophthiriidae 

Antarctophthirus tricechi (Boheman 1865) 176 

Echinophthirius horridus (Olfers 1816) 

Dermaptera 

_ 

_ 

Echinophthirius groenlandicus 

Becher 

Forficulidae 

Forficula auricularia L 1758 

Dictyoptera 

_ 

_ 

Blattellidae 

Blattella germanica (L. 1767) 

Ephemeroptera 

_ 

_ 

Leptophlebiidae 

Leptophlebia vespertina (L. 1758) 173 

110 176 


Hemiptera 

_ 

_ 


Aphidoidea 

Sitobion calvulum (Ossiannilsson 1958) 

Acyrthosiphon calvulum 

Ossiannilsson 1958 

139 245 411 419 508 507 

Acyrthosiphon svalbardicum Heikinheimo 1968 139 150 156 294 295 296 297 298 299 300 301 334 344 

347 348 387 408 411 415 420 421 454 509 514 552 569 

592 594 599 602 

Cinara abieticola (Cholodkovsky 1899) 

?Cinara piceae (Panzer 1801) ?Dilachnus piceae (nec Walker) 

105 139 

Pemphigus groenlandicus (Rübsamer 1898) 139 145 

Phthiraptera 

_ 

_ 

Amblycera 

Ancistrona vagelli (J.C. Fabricius 1787) 224 


420


Austromenopon brevifimbriatum (Piaget 1880) 224 

Austromenopon corporosum (Kellogg and Kuwana 1901) 224 

Austromenopon fuscofasciatum (Piaget 1880) 

Menopon fuscofasciatum Piaget 

Austromenopon lutescens (Burmeister 1838) 

Menopon lutescens Burmeister 

Austromenopon merguli Timmermann 1954 224 

Austromenopon uriae Timmermann 1954 224 

126 224 

110 126 224 319 

Trinoton anserinum (J.C. Fabricius 1805) 126 224 308 

Ischnocera 

Anaticola anseris (L. 126 224 



Anaticola crassicornis rubromaculatus (Rudow 1869) 224 

Carduiceps zonarius (Nitzsch [in Giebel],1866) 

Carduiceps complexivus (Kellogg 

and Chapman) 

Goniodes lagopi (L. 1758) 224 

Lagopoecus pallidovittatus (Grube 1851) 

Nirmus quadrulatus Nitzsch Lagopoecus cameratus (Nitzsch and 

Lyonet) 

Lunaceps neiris Timmermann 1954 

? Degeegeriella actophilus (Kellogg 

and Chapman 1899) 

126 224 

126 224 

126 224 319 

Lunaceps phaeopi (Denny 1842) 126 224 

Ornithobius hexophtalmus (Nitzsch 1871) 126 224 

Perineus nigrolimbatus (Giebel 1874) 

Esthiopterum nigrolimatum (Giebel) Lipeurus mutabilis Piaget 

110 126 124 224 319 



Quadraceps aethereus (Giebel 1874) 

Cummingsiella aetherea klatti 

(Timmermann 1954) 

Mjöberginirmus klatti (Timmermann) 

Quadraceps hiaticulae (O. Fabricius 1780) 

Chadraceps hiaticulae (O. 

Fabricius) 1780 

Quadraceps normifer (Grube 1851) 

Cummingsiella normifer (Grube 

1851) 

Quadraceps obliquus (Mjöberg 1910) 

Cummingsiella obliqua aquilonis 

Timmermann 1974 

Cummingsiella klatti (Timmerman 

1954) 

Cummingsiella hiaticulae (O. 

Fabricius 1780) 

Quadraceps ornatus striolatus (Nitzsch [in Giebel], 1866) 

Cummingsiella ornatus (Grube 

1851) 

Quadraceps phalaropi (Denny 1842) 

Cummingsiella phalaropi (Denny 

1842) 

224 

124 224 

224 

224 

224 

224 



Quadraceps strepsilaris (Denny 1842) 

Cummingsiella strepsilaris (Denny 

1842) 

Saemundssonia (Saemundssonia) calva (Kellogg 1896) 124 224 

Saemundssonia (Saemundssonia) cephalus (Denny 1842) 126 224 

Saemundssonia (Saemundssonia) grylle (O. Fabricius 1780) 126 224 

Saemundssonia (Saemundssonia) inexspectata Timmermann 

1951 

Saemundssonia (Saemundssonia) lari (O. Fabricius 1780) 

Docophorus gonothorax (Giebel) 

Docophorus lari Denny 

Philopterus sp. 

Saemundssonia (Saemundssonia) lockleyi Clay 1949 

? Saemundssonia sternae (L.) 

224 

224 

110 126 124 224 

126 224 

Saemundssonia (Saemundssonia) merguli (Denny 1842) 126 124 224 



Saemundssonia (Saemundssonia) occidentalis (Kellogg 1896) 124 224 

Saemundssonia (Saemundssonia) scolopacisphaeopodis 

(Schrank 1803) 

Saemundssonia (Saemundssonia) tringae tringae (O. Fabricius 

1780) 

Philopterus fusiformis (Denny) 

Saemundssonia (Saemundssonia) tringae variabilis (Denny 1842) 

Saemundssonia (Saemundssonia) tringae lobata Martens 1974 224 

Thysanoptera 

_ 

_ 

Thripidae 

Aptinothrips rufus Haliday 1836 410 

126 224 

124 224 319 


Crustacea 

Branchiopoda 

Cladocera 

_ 

_ 


Bosminidae 

Bosmina longirostris Müller 1785 

Eubosmina longispina (Müller 1785) 

Bosmina longispina Müller Bosmina obstusirostris Sars 

Chydoridae 

Acroperus harpae (Baird 1835) 19 20 165 170 466 

Alona guttata Sars 1862 165 173 466 

Chydorus sphaericus (Müller 1785) 19 

559 

20 130 165 170 171 173 227 305 316 318 319 466 

Daphniidae 



Daphnia pulex 

Species complex 

Daphnia pulicaria 

Daphnia longiremis 

Daphnia (Hyalodaphnia) umbra 

Daphnia middendorffiana 

Daphnia tenebrosa 

Daphnia longispina 

7 9 19 20 64 92 130 144 148 149 165 170 171 

173 191 195 196 202 218 225 227 305 316 318 319 337 

338 339 356 357 395 390 391 401 402 403 404 406 435 

466 491 491 527 540 559 

225 

Macrothricidae 

Macrothrix hirsuticornis Norman and Brady 1867 130 

559 

165 170 171 173 226 275 276 305 311 319 466 556 

Macrothrix hirsuticornis arctica (Sars 1890) 

Ctenopoda 

_ 

_ 

Macrothrix arctica Sars 

20 19 227 318 319 

Sididae 

Sida crystallina (Müller 1776) 19 20 173 

Notostraca 

_ 

_ 

Triopsidae 

Lepidurus arcticus (Pallas 1793) 19 20 130 171 173 188 195 196 305 318 319 391 405 

466 479 512 510 536 


Copepoda 

Calanoida 

_ 

_ 


Centropagidae 

Limnocalanus macrurus Sars 1863 173 311 

Temoridae 

Eurytemora raboti Richard 1897 9 130 171 305 318 319 527 559 

Cyclopoida 

_ 

_ 

Cyclopidae 

Acanthocyclops gigas (Claus 1857) 

Megacyclops gigas (Claus) Cyclops gigas Claus 

Cyclops abyssorum Sars 1863 

Cyclops strenuus Fischer Cyclops strenuus medianus 

19 20 197 319 305 

9 

559 

19 20 103 130 173 195 197 318 319 390 391 491 

Cyclops vicinus Ulianine 1875 19 20 103 197 305 319 317 



Diacyclops crassicaudis (Sars 1863) 

Harpacticoida 

_ 

_ 

Cyclops crassicaudis Sars 

Canthocamptidae 

Maraenobiotus brucei brucei (Richard 1898) 

Maraenobiotus brucei (Richard 

1898) 

130 171 173 197 311 318 

311 536 

Harpacticidae 

Harpacticus uniremis Kröyer 1845 305 319 

Tachidiidae 

Tachidius discipes Giesbrecht 1882 

Tachidius brevicornis Lilljeborg ?Tachidius spitsbergensis Olofsson 

Siphonostomatoida 

_ 

_ 

305 315 318 

Lernaeopodidae 

Salmincola edwardsii Olsson 1869 184 277 



Salmincola carpionis (Krøyer 1837) 396 

Malacostraca 

Amphipoda 

_ 

_ 

Gammaridae 

Gammaracanthus lacustris (Sars 1867) 396 

Gammaracanthus loricatus (Sars 1896) 135 

Mysidacea 

_ 

_ 

Mysidae 

Mysis relicta Loven 1861 

Mysis oculata (Fabricus) var. relicta 

305 496 


Ostracoda 

Podocopida 

_ 

_ 


Candonidae 

Candona candida Müller 1776 20 319 

Eucandona rectangulata (Alm 1914) 

Candona rectangulata Alm 

Cyprididae 

Cyclocypris ovum (Jurine 1820) 319 

Eucypris arctica Olofsson 1918 305 319 

Eucypris glacialis (Sars 1890) 

Prionocypris glacialis (Sars) 

Heterocypris fretensis (Brady and Robertson 1870) 136 

136 305 311 319 318 

19 20 136 319 305 496 



Tonnacypris glacialis (G.O.Sars, 1890) 524 

Cytherideidae 

Cytherissa lacustris (Sars 1863) 311 491 

Limnocytheridae 

Limnocythere inopinata (Baird 1843) 136 


To be classified 

_ 

_ 

_ 

_ 

_ 


Cestoda 

Nadejdolepsis nitida 506 

Aploparakis leonovi 506 

Aploparakis crassipenis 506 


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Checklist of the Terrestrial and freshwater invertebrate fauna ... - UNIS

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