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Untitled - The Alfred Russel Wallace Website

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ANTS AND PLANT-STRUCTURE 407<br />

[When I compare these and similar instances<br />

with the Pitchers of the Nepenthes, in which (as<br />

I<br />

learn from the accounts of travellers) ants as well as<br />

water are nearly always found,<br />

I cannot doubt that<br />

those curious appendages have attained their actual<br />

dimensions through the deepening 'and widening<br />

which they have undergone from ants through<br />

untold ages.]<br />

We have a curious example, in the genus<br />

Cinchona, of the supposed correlation of a minute<br />

structural peculiarity with chemical and medical<br />

properties. Eminent botanists, 'such as Weddell<br />

and Karsten, who have studied that ^enus in its<br />

c><br />

native forests, have thought they had found a char-<br />

acter in the leaves always associated with a bark<br />

rich in alkaloids, viz. the presence of a small pit or<br />

scrobicule in the axil of each vein on the underside<br />

of the leaf. But when good specimens of C. sitc-<br />

cirubra, the richest of all the barks in alkaloids,<br />

came to be examined, the leaves were found entirely<br />

destitute of scrobicules ! See now how this comes<br />

about. <strong>The</strong> leaves of the Hill Barks those, namely,<br />

that grow at an elevation of 8000 feet and upwards<br />

are liable to be infested by a small mite which<br />

nestles in the scrobicules has caused them, in fact-<br />

its remote ancestors having at first sheltered in the<br />

vein-axils but C. succirubra ; grows always below<br />

that elevation indeed, as low down as 2400 to 6000<br />

feet and is the only quinine-producing Cinchona<br />

that descends so low, the other species of Cinchona<br />

that grow at a low elevation having all medically<br />

worthless bark. But as all these species, C. suc-<br />

cirubra included, are equally destitute of scrobiculate<br />

leaves and of mites, the reasonable inference is that

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