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Untitled - The Alfred Russel Wallace Website

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xxa ANTS AND PLANT-STRUCTURE 397<br />

its allies. <strong>The</strong> opposite leaves, 9 inches long, are<br />

oblong-oval, obtuse with a short apiculus, near the<br />

base abruptly panduriforni, and bearing a small ant-<br />

sac on the midrib. All the other known species of<br />

this large genus have non-sacciferous leaves.<br />

In all the plants I have seen bearing sacs on the<br />

leaves, to whatever order they belong, it is remark-<br />

able that the pubescence consists of long hairs<br />

having a tubercular base ; and<br />

although I do not<br />

see what connection that peculiarity can have with<br />

the ants' choice of a habitation, it is probable they<br />

find some advantage in it.<br />

2. Of Inflated Petioles<br />

A true swelling of the petiole, inhabited by ants,<br />

and (as I believe) owing its existence to their<br />

agency, I have seen only in two genera of Leguminose<br />

Coesalpiniese,<br />

viz. Tachigalia and Sclero-<br />

lobium. <strong>The</strong> Tachigalia^ are low-growing riparial<br />

trees, of black-water rivers, and have pinnate, often<br />

silky foliage, and small, yellow, sweet-smelling, nearly<br />

regular flowers disposed in<br />

panicles. All have<br />

trigonous petioles, which are mostly dilated at the<br />

base into a fusiform sac tenanted by ants. T. caripcs,<br />

sp. n., grows abundantly on the banks, and on<br />

inundated islands, of the Uaupcs. It is a spreading<br />

tree of 30 feet, and has the ramuli, petioles, and<br />

leaves clad with a fine, close, silky pubescence.<br />

<strong>The</strong> sacs of the petiole are inhabited by small black<br />

ants, whose entrance is by a little hole on the<br />

underside of the sac. T. ptychophysca, sp. n., grows<br />

in moist sandy caatingas by the same river, and has<br />

a similar sac on the petiole.

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