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Untitled - The Alfred Russel Wallace Website

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ANTS AND PLANT-STRUCTURE 389<br />

which is adjacent to and inundated by the rivers<br />

and lakes, but sometimes deep in the virgin forest,<br />

wherever the land is so low that the water of rains<br />

may accumulate thereon to a slight depth. All the<br />

species have the unmistakable aspect of their order<br />

-the ribbed opposite leaves, the polypetalous flowers<br />

with beaked porose anthers, etc. ; but<br />

they are dis-<br />

tinguished at sight from most others of the order<br />

by the large, thin, lanceolate or ovate acuminate,<br />

leaves, very sparsely set with long hairs, and having<br />

a hollow sac or a pair of sacs at the base either of<br />

all the leaves, or (more frequently) of only one of<br />

each pair when that one is much larger than the<br />

other. <strong>The</strong> leaves in the majority of the species<br />

have but three ribs ; a few species, however, have<br />

five- or even seven-ribbed leaves ; but, in all, the<br />

origin of the innermost pair of ribs is an inch or so<br />

up the midrib from the base of the leaf; and it is<br />

this portion of the leaf, from the insertion of the<br />

inner ribs downwards, which is occupied by the sac.<br />

<strong>The</strong> latter sometimes takes up only a part of the<br />

breadth of the leaf, when it is technically considered<br />

to be seated on the leaf ; (Epiphysca) in other<br />

cases the sac in its lower half absorbs the whole<br />

breadth of the leaf, when it seems to be seated half<br />

on the leaf, half on the petiole (Anaphysca) ;<br />

or,<br />

lastly, throughout its length it absorbs the whole<br />

breadth of the leaf, and then seems seated entirely<br />

on the petiole (Hypophysca). That it is really<br />

formed in all cases at the expense of the lamina,<br />

and not of the petiole, is proved by the occasional<br />

occurrence of imperfectly -developed sacs in the<br />

hypophyscous form, bordered by a narrow wing continuous<br />

with the leaf, and Diviner to the latter a

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