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16. Radiolaria - Deep Sea Drilling Project

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M. G. PETRUSHEVSKAYA, G. E. KOZLOVA<br />

Genus LIPMANELLA Loeblich and Tappan,<br />

emend. Petruskevskaya<br />

Lipmanella Loeblich and Tappan, 1961, p. 119. = Dictyoceras<br />

Haeckel, 1860, p. 333; 1887, p. 1324; Campbell, 1954, p. 130;<br />

Petrushevskaya, 1971b, pp. 198, 220. Type species Lithornithium<br />

dictyoceras Haeckel (Haeckel, 1862, pi. 8, figs. 1-5).<br />

The cephalis thick-walled, divided (as a rule) into upper and<br />

lower parts by means of the arches ap, as in Squinabollum and in<br />

Dicolocapsa. The thorax is larger than the cephalis. The abdomen<br />

may be separated distinctly from the thorax or it may be united<br />

with the thorax in a single segment. The spines D, LI and Lr form<br />

external feet, which are rather weak. They go laterally from the<br />

thorax. In the upper walls of the thorax are three ribs formed by<br />

these spines. The spine A forms the apical horn. The mouth of the<br />

shell is very often open.<br />

Cretaceous?-Quaternary.<br />

Remark: The taxonomical position of the genus is doubtful-maybe<br />

it is really pterocoryid.<br />

Lipmanella (?) sp. aff. Lithomelissa? amazon Foreman<br />

(Plate 37, Figure 1)<br />

^.Lithomelissa amazon Foreman, 1968, p. 26, pi. 4, fig. 1.<br />

The division of the cephalis into lower and upper parts is not<br />

developed. In typical Lithomelissa amazon, the cephalis is smoother.<br />

Paleogene.<br />

Lipmanella (?) sp. M<br />

(Plate 37, Figure 2)<br />

The thorax, the spines and the pores are very much the same as<br />

in Lipmanella (?) sp. aff. Lithomelissa amazon. The difference is in<br />

the smaller dimensions and in the cephalis being constricted at its<br />

base, with pronounced subdivision into upper and lower parts.<br />

Miocene.<br />

Lipmanella sp. O<br />

(Plate 37, Figures 4,5)<br />

Differs from Lipmanella (?) sp. M by the distinct subdivision of<br />

the skeleton into thorax and abdomen.<br />

This species is somewhat similar to Dictyoceras insectum<br />

Haeckel (1887, pi. 71, fig. 6), and there is no doubt in the generic<br />

identification of this species (Dictyoceras = Lipmanella). It is also<br />

somewhat similar to Pterocorys columbo Haeckel (1887, pi. 71, fig.<br />

2), but differs in the construction of the cephalis. Some specimens<br />

are similar to Pterocyrtidius barbadense (Ehrenberg) (see Plate 27,<br />

Figure 18, 19) in the proportions of the test.<br />

Miocene.<br />

Lipmanella (?) dogieli (Petrushevskaya)<br />

(Plate 37, Figure 10)<br />

Sethoconus (?) dogieli Petrushevskaya, 1967, p. 95, pi. 53, figs. 1,2;<br />

1971b, pi. 110, fig. 2.<br />

The number of pores is less than in Lipmanella sp. C-only about<br />

7 pores on a half equator. Pores are irregular and of different sizes.<br />

There is no subdivision into thorax and abdomen.<br />

Pliocene-Quaternary.<br />

Remark: This species is very similar also to the Miocene species<br />

described as Lithomelissa campanulaeformis by Campbell and<br />

Clark (1945, p. 44, pi. 6, fig. 1), see Plate 37, Figure 11 in the<br />

present report. Lipmanella (?) dogieli differs from L. campanulaeformis<br />

by smaller dimensions. Perhaps it represents by itself only<br />

Quaternary subgenus of the Miocene species L. campanulaeformis.<br />

Subfamily Group E<br />

To include genera without a wide velum (characteristic for<br />

Neosciadiocapsidae). In place of the velum there is a long, nearly<br />

cylindrical abdomen. Abdominal pores are much larger than<br />

thoracic pores, and therefore the thorax and abdomen seem to be<br />

built of different kinds of skeletal meshwork. Cephalis of the same<br />

kind as in Neosciadiocapsidae, with the arches ap going in the upper<br />

walls. Cephalis conical, wide at the base. The apex gradually turns<br />

into a conical apical horn. Lampterium is the most typical genus of<br />

the group. Whether this group should be placed among Pterocoryidae<br />

or among Neosciadiocapsidae is questionable.<br />

542<br />

Genus THYRSOCYRTIS Ehrenberg<br />

Thyrsocyrtis Ehrenberg, 1847b, p. 54; Haeckel, 1887, p. 1350;<br />

Campbell, 1954, p. 130; Riedel and Sanfilippo, 1970, p. 525.<br />

Type species Thyrsocyrtis rhizodon Ehrenberg, 1873 (1875, pi.<br />

12, fig. 1).<br />

Podocyrtarium Haeckel, 1887, p. 1337; Campbell, 1954, p. 130.<br />

Type species Podocyrtis tripodiscus Haeckel, (1887, pi. 72,<br />

fig. 4).<br />

Podocyrtonium Haeckel, 1887, p. 1347, Campbell, 1954, p. 130.<br />

Podocyrtis pedicellaria Haeckel (1887, pi. 72, fig. 8).<br />

Differs from Lampterium by distinct longitudinal rows of pores<br />

not only on the thorax, but also on the abdomen. Differs from<br />

Podocyrtis 1) by the abdomen being larger than the thorax, and<br />

2) by the difference in the size between thoracic and abdominal<br />

pores.<br />

Tertiary.<br />

Thyrsocyrtis rhizodon Ehrenberg<br />

Thyrsocyrtis rhizodon Ehrenberg, 1873, p. 262; 1875, pi. 12, fig. 1;<br />

Riedel and Sanfilippo, in press, pi. 3C, fig. 6.<br />

Eocene.<br />

Thyrsocyrtis tetracantha (Ehrenberg)<br />

Podocyrtis tetracantha Ehrenberg, 1873, p. 254; 1875, pi. 13, fig. 2.<br />

Thyrsocyrtis tetracantha (Ehrenberg) Riedel and Sanfilippo, 1970,<br />

p.527.<br />

Eocene.<br />

Thyrsocyrtis triacantha (Ehrenberg)<br />

(Plate 32, Figure 9; Plate 34, Figure 6)<br />

Podocyrtis triacantha Ehrenberg, 1873, p. 254; 1875, pi. 13, fig. 4.<br />

Thyrsocyrtis triacantha (Ehrenberg) Riedel and Sanfilippo, 1970, p.<br />

526, pi. 8, figs. 2, 3; Moore, pi. 4, fig. 2.<br />

Eo cene-Oligocene ?<br />

Thyrsocyrtis argulus (Ehrenberg)<br />

(Plate 32, Figure 8)<br />

Podocyrtis argulus Ehrenberg, 1873, p. 248; 1875, pi. 16, fig. 2.<br />

Thyrsocyrtis hirsuta hirsuta (Krasheninnikov) Riedel and Sanfilippo,<br />

1970, p. 7, figs. 8,9.<br />

Eocene.<br />

Thyrsocyrtis sp. E<br />

(Plate 32, Figure 11)<br />

IPodocyrtis pedicellaria Haeckel, 1887, p. 1347, pi. 72, fig. 8.<br />

Wodocyrtis sinuosa (?) in Riedel and Sanfilippo, 1970, p. 534, pi.<br />

11, figs. 3,4.<br />

Eocene.<br />

Podocyrtis (?) or Thyrsocyrtis (?) mitra (Ehrenberg)<br />

Podocyrtis mitra Ehrenberg, 1854, pi. 36, fig. B 20; 1873, p. 251;<br />

non Ehrenberg, 1875, pi. 15, fig. 4.<br />

Podocyrtis (Lampterium) mitra (Ehrenberg) Riedel and Sanfilippo,<br />

1970, p. 534, pi. 11, figs. 5, 6; Moore, in press, pi. 3, fig. 4.<br />

Eocene.<br />

Thyrsocyrtis ? bromia Ehrenberg<br />

Thyrsocyrtis bromia Ehrenberg, 1873, p. 260; 1875, pi. 12, fig. 2;<br />

Riedel and Sanfilippo, 1970, p. 526; Riedel and Sanfilippo, in<br />

press, pi. 8, fig. 6, Moore, in press, pi. 5, figs. 1-3.<br />

Cephalis and thorax very similar to Cyclamp terium species, but<br />

abdomen is smaller.<br />

Eocene.<br />

Genus LOPHOCYRTIS Haeckel<br />

Lophocyrtis Haeckel, 1887, p. 1410; Campbell, 1954, p. 134;<br />

Riedel and Sanfilippo, 1970, p. 529. Type species Eucyrtidium<br />

stephanophorum Ehrenberg (1873, p. 223; 1875, pi. 8,fig. 14).<br />

Differs from Thyrsocyrtis mainly by the abdomen being<br />

narrower.<br />

Lophocyrtis? jacchia (Ehrenberg) group<br />

(Plate 28, Figure 21)<br />

Thyrsocyrtis jacchia Ehrenberg, 1873, p. 261; 1875, pi. 12, fig. 7.<br />

Thyrsocyrtis dionisia Ehrenberg, 1873, p. 260; 1875, pi. 12, fig. 5.

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