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16. Radiolaria - Deep Sea Drilling Project

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M. G. PETRUSHEVSKAYA, G. E. KOZLOVA<br />

Stylodictya inaequalispina Clark and Campbell<br />

(Plate 18, Figure 8)<br />

Stylodictya inaequalispina Clark and Campbell, 1942, p. 45, pi. 3,<br />

fig. 5.<br />

Disc diameter 150-190µ; width of rings 12µ, pore diameter 4-6µ<br />

(in center 2µ), length of spines up to 50µ.<br />

Eocene-Oligocene.<br />

Stylodictya rosella, Kozlova sp. n.<br />

(Plate 18, Figure 9)<br />

Stylodictya sp. Clark and Campbell, 1945, p. 24, pi. 3, fig. 19.<br />

Central chamber is surrounded by 9-11 slightly curved (scalloped)<br />

annular systems, each system doubling in size beginning in<br />

the center to the edge of the disc; pores are found, 2 or 3 on each<br />

ring, approximately of the same size; there are up to twenty-four<br />

inner spines of uniform thickness; at the rim of the disc they appear<br />

either as thin spicules or as massive flat teeth.<br />

Disc is very thin in its central part and becomes very thick at the<br />

rim (the last 5 to 6 rings).<br />

Disc diameter 250-270µ, width of rings 7-16µ, pore diameter<br />

3-4µ; diameter of the first system 24µ.<br />

It differs from S. targaeformis (see below) by larger dimensions,<br />

the thickness along its rim, and by a greater number of spines.<br />

Eocene-Oligocene.<br />

Stylodictya targaeformis (Clark and Campbell)<br />

(Plate 18, Figure 10)<br />

Staurodictya targaeformis Clark and Campbell, 1942, p. 43, pi. 3,<br />

fig. 6.<br />

Specimens from Atlantic have eight external spines.<br />

Disc diameter 170-200µ; width of rings 5-10µ; pore diameter<br />

2-5µ, length of spines 40µ; diameter of the first system 30µ.<br />

Eocene-Oligocene.<br />

Stylodictya orbiculata (Haeckel)<br />

(Plate 18, Figure 7)<br />

Trematodiscus orbiculatus Haeckel, 1862, pi. 28, fig. 2.<br />

Porodiscus orbiculatus Haeckel, 1887, p. 492.<br />

Disc diameter 125-140µ, width of rings 7-15µ;pore diameter 5µ.<br />

Eocene-Oligocene.<br />

Stylodictya aculeata (Jörgensen)<br />

(Plate 18, Figure 6)<br />

Stylodictya aculeata Jörgensen, 1905, p. 119, pi. 10, fig. 41;<br />

Petrushevskaya, 1967, p. 35, pi. 11, figs. 1-3.<br />

Quaternary.<br />

CIRCODISCUS Kozlova, new genus<br />

Porodiscus Haeckel, 1881, p. 491, (part.); Ommatodiscus Stöhr,<br />

1880, p. 115 (part.); Flustrella Campbell, 1954, p. 89 (part.)<br />

Type species Trematodiscus microporus Stöhr, 1880, p. 108, pi.<br />

4, fig. 17.<br />

Slightly convex skeleton with round or oval outline. Consists of<br />

frontal, sagittal and equatorial girdles, their wings connected to<br />

form regular round or oval rings.<br />

First system is of the Trizonium type {in sensu Haeckel), its<br />

dimension 30-40µ, with fully developed girdles; in the rest of<br />

them, only the equatorial girdle is fully developed; saggital and<br />

frontal girdles terminate at the point of the junction with the ring of<br />

the previous system; distance between (width of rings) is considerably<br />

larger than the diameter of the central chamber. Spines (four<br />

main and four secondary) pass inside of the skeleton. Rim is<br />

smooth.<br />

It differs from Plectodiscus gen. nov., by the character of the<br />

connection of its wings and accordingly by the structure of the first<br />

system.<br />

The name is derived from Greek "Circos" (circle) and "Discos"<br />

(disk), masculine.<br />

Eocene-Recent.<br />

Circodiscus microporus (Stöhr)<br />

(Plate 19, Figure 1-7)<br />

Trematodiscus microporus Stöhr, 1880, p. 108, pi. 4, fig. 17.<br />

Porodiscus microporus Stöhr, Haeckel, 1887, p. 493.<br />

526<br />

Skeleton is oval, flat, consisting of 3 or 4 comparatively wide,<br />

oval, annular girdles; pores are even, circular, 5 or 6 on each ring.<br />

Inner spines (8?) do not extend outside; rim of the disc seldom has a<br />

smooth, clearly outlined tapered edge; more often it is "torn" and<br />

disconnected.<br />

Disc diameter with four systems 180-200µ width of rings<br />

25-35µ, pore diameter 5-8µ.<br />

Miocene-Quaternary.<br />

Circodiscus sp.<br />

(Plate 19, Figure 8)<br />

From C. microporus it differs by having wider rings and smaller<br />

pores.<br />

Disc diameter (with three systems) 160µ, width of rings 3040µ,<br />

pore diameter 3-4µ.<br />

Miocene.<br />

Genus PLECTODISCUS Kozlova, new genus<br />

Discospira Haeckel, 1862 (part.); Porodiscus Haeckel, 1882, p. 492<br />

(part.); Flustrella Campbell 1954, p. 89 (part.). Type species<br />

Porodiscus circularis Clark and Campbell, 1945, p. 42, pi. 11,<br />

figs. 2, 6, 10.<br />

Thin, biconvex skeleton, with oval, less frequently circular<br />

outline, consisting of frontal, equatorial and sagittal girdles. The<br />

closed-spiral structure of girdles is of the Spironium type: wings of<br />

the same type of girdle form a one-half turn of the spiral, and at the<br />

point of their juncture overlap each other. First system has fully<br />

developed girdles, its dimensions are 40 × 50µ; in the rest of the<br />

girdles, only the equatorial girdle is fully developed, both saggital<br />

and frontal girdles terminating at the point of their junction with<br />

the previous system. Distance between rings is greater than is the<br />

diameter of the central chamber. Inner spines are indistinct; rim of<br />

the disc is smooth or is covered by numerous short spines.<br />

Depending on the degree of inclination of the plane of the<br />

girdles in relation to the main axes of the skeleton, the equatorial<br />

girdles appear from the surface (rest of girdles in corresponding<br />

cross-section) in the form of rings, as a single or double spiral, or in<br />

the form of rings connected by zigzag radial lines.<br />

The name is derived from Greek "plecta" (interlacing) and<br />

"Discos" (disk), masculine.<br />

Paleocene-Oligocene.<br />

Plectodiscus circularis (Clark and Campbell)<br />

(Plate 19, Figures 9-12)<br />

Porodiscus circularis Clark and Campbell, 1942, p. 42, pi. 11, figs. 2,<br />

6, 10.<br />

Porodiscus uralicus Lipman, 1960, p. 86, pi. XI, figs. 9-11;<br />

Porodiscus durus Moksyakova, p. 146, pi. 2, fig. 8.<br />

Skeleton is oval, less frequently circular in outline, consisting of<br />

4 to 6 systems, the width of which increases slightly from the center<br />

toward the rim of the disc; all possible varieties of the outer forms<br />

of the girdle are represented here: the enclosed-spiral (Plate-Figure<br />

12), annular (Plate-Figures 9, 10) and spiral; pores are round and of<br />

uniform size, four or five on the width of each system; disc rim is<br />

tapered; main and secondary radial spines are of uniform thickness<br />

and appear at the rim of the disc in the form of (16?) tiny spicules.<br />

Disc diameter 120-25Oµ, distance between systems 20-35µ, pore<br />

diameter 6-9µ.<br />

Paleocene?-Eocene. Paleocene forms have a coarser and larger<br />

skeleton.<br />

Plectodiscus bergontianus (Carnevale)<br />

(Plate 19, Figure 13)<br />

Porodiscus bergontianus Carnevale, 1908, p. 23, pi. 4, fig. 2.<br />

Porodiscus squinaboli Carnevale, 1908, p. 23, pi. 4, fig. 3.<br />

Disc diameter 230-240µ, width of rings 15-20µ, pore diameter<br />

5-7µ.<br />

This form differs from P. circularis by the greater number of<br />

systems and their smaller width.<br />

Eocene.<br />

Genus OMMATOCAMPE Ehrenberg<br />

Ommatocampe Ehrenberg, 1860, p. 852; Haeckel, 1887, p. 392;<br />

Campbell, 1954 p. 76. Type species Ommatocampe polyarthra<br />

Ehrenberg, 1872b, pi. 6, fig. 9.

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