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OFR 151.pdf - CRC LEME

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troughs in the Bass and Gippsland Basins. Onset of seafloor spreading in the Tasman Sea at<br />

about 80 Ma led to the separation of New Zealand and a number of other continental<br />

fragments from the palaeo-eastern margin of Australia. Associated events included igneous<br />

activity in southeastern Queensland and eastern New South Wales. An extensive river system<br />

drained into the Indian Ocean via the Officer and Canning Basins. Elsewhere deep<br />

weathering continued over much the continent.<br />

6.6.2 Palaeobotany<br />

Forests and woodlands remained dominated by gymnosperms although the diversity and<br />

relative abundance of angiosperms continued to increase. Whether these angiosperms were<br />

shrubs or trees is uncertain. In palynostratigraphic terms, the most prominent clade to migrate<br />

into Australia at this time (at the Santonian/Campanian boundary) were ancestral species of<br />

Nothofagus, represented by the fossil pollen species Nothofagidites endurus and N. senectus<br />

(Dettmann et al. 1990). Fossil pollen representing the four extant subgenera of Nothofagus<br />

(Hill and Read 1991) are not recorded earlier than the Late Campanian. Other newly arrived<br />

taxa include horsetails (Ephedra), palms (Longapertites) and a fossil member of the<br />

Didymelaceae, which now is confined to angiosperm rainforest in Madagascar (Schizocolpus<br />

marlinensis).<br />

6.6.3 Palaeoclimates<br />

Assuming that ancestral Nothofagus has similar preferences to modern Nothofagus,<br />

differences in relative abundance of Nothofagidites indicates strong contrasts in rainfall and<br />

mean annual temperatures between the palaeo-northern and southern margins. The presence<br />

of palms (Palmae) is assumed to indicate very warm to hot conditions.<br />

Palaeo-northern Australia<br />

Podocarp-dominated Austral Conifer Forest and fern heath remained the dominant vegetation<br />

types in coastal regions throughout the Early Campanian. Conditions are likely to have been<br />

wet to very wet (humid-perhumid) and very warm (upper mesotherm) based on the presence<br />

of palms. Rare taxa in the offshore sequences include two probable temperate rainforest taxa,<br />

Lophosoria and Nothofagus (Nothofagidites sp. cf. N. senectus), which may imply conditions<br />

away from the coast were cooler with more uniform rainfall than in the Turonian-Santonian.<br />

Palaeo-central Australia<br />

No fossil evidence is available and it is uncertain whether temperate rainforest angiosperms<br />

such as Nothofagus were present in central Australia at this time.<br />

Palaeo-southern Australia<br />

Austral Conifer Forest and fern heath remained the dominant vegetation types in the southern<br />

margin basins although araucarians are less common than during the Turonian-Santonian,<br />

possibly because of different depositional contexts (reduced Neves Effect), and ancestral<br />

species of Nothofagus (Nothofagidites endurus, N. senectus) became frequent to common at<br />

many sites in the late Early Campanian. The latter is consistent with increasingly wet<br />

(perhumid) and/or more uniform conditions, although mean annual temperatures are likely to<br />

have remained within the microtherm range due to the high palaeolatitude (~70 0 S).<br />

76

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