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OFR 151.pdf - CRC LEME

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1.3 Flora, vegetation and climate<br />

1.3.1 Flora<br />

An important distinction exists between flora and vegetation. A flora comprises the plant<br />

species present in an area irrespective of their relative abundance or life form. Vegetation<br />

denotes the abundance and structural arrangement of the taller or more common species.<br />

Plant community is used to denote relatively stable associations between particular species,<br />

which in combination make up particular vegetation types. Species making up the major<br />

vegetation types in Australia are given in Specht (1970, 1982).<br />

1.3.2 Vegetation<br />

Vegetation has a staggering potential to alter global and regional environments in the shortterm<br />

and over geological time. For example, on average plants contain less than 0.5 g carbon<br />

cm 3 and form a ‘veneer’ that is less than 100 m thick between the 100 km deep lithosphere<br />

and 10 km high atmosphere. Nevertheless, the global vegetation manages to cycle 60<br />

gigatonnes (60 x 10 15 g) of carbon between the lithosphere, biosphere and atmosphere each<br />

year (Naeem 1999).<br />

Similarly, methane produced by the bacterial decomposition of organic matter in wetlands has<br />

been implicated as a major factor in global warming. A brief but intense period of global<br />

warming and massive perturbation of the global carbon cycle during the Paleocene-Eocene<br />

transition (Paleocene-Eocene thermal maximum) almost certainly was due to the catastrophic<br />

release over ~20 ka of methane, which previously had been ‘frozen’ as hydrates in sea floor<br />

sediments (Rohl et al. 2000).<br />

1.3.3 Biomes<br />

Biomes are defined as broad stable ecosystems of plants and animals, which occupy climatic<br />

regions defined by unique combinations of precipitation and temperature, and seasonal<br />

variation in these parameters. Plant biomes are broad vegetation classes defined by<br />

combinations of dominant physiognomic character (life form, leaf-form, phrenology).<br />

Terrestrial and most aquatic ecosystems are underpinned by plants whose life-forms/size<br />

(trees, shrubs, herbs), growth rates and distributions are shaped not only by the prevailing<br />

climate but also by soil fertility, drainage and moisture status (edaphic factors).<br />

Global biomes<br />

The strong association of particular life forms with particular climatic regimes allows plant<br />

biomes to be defined using a combination of climatic, physiognomic and physiological<br />

characters (Christopherson 1997). Because of functional convergence between unrelated<br />

plants growing under the same physical environment, it is possible to use ecophysiological<br />

data to compare or combine data from regions with taxonomically different floras (Prentice et<br />

al. 1992).<br />

Eleven terrestrial plant biomes can be recognised globally (Table 1). Most biomes are named<br />

after the tallest/most abundant (dominant) plant stratum present, i.e. the one most exposed to<br />

climate. They are rainforest, savannah (grasslands with scattered trees), grassland (grasslands<br />

lacking woody species), shrublands, desert and tundra. Each is characterised by a distinctive<br />

flora with equally distinctive morphological and ecophysiological adaptations. The transition<br />

between biomes is recognisable by changes in the physiognomic character of the vegetation<br />

as a whole, in the relative proportions of the component plant communities, or in the relative<br />

abundance of the dominant plant species.<br />

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