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OFR 151.pdf - CRC LEME

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Nothofagus-gymnosperm temperate rainforest within (Late Miocene, Late Pliocene)<br />

predominantly wet sclerophyll forest types.<br />

Inferred climate<br />

The expansion of Nothofagus (Lophozonia) spp. is likely to be due to uniformly wet<br />

(perhumid) and cool (upper microtherm) conditions on the upper slopes of the Southern<br />

Highlands during an interval (Early Pliocene) when other sites in southern Australia<br />

experienced increasing warmth. The most plausible explanation is that warm sea surface<br />

temperatures and high relative sea levels resulted in increased orographic cloudiness, and<br />

therefore effectively cooler wetter conditions, over the Southeastern Highlands.<br />

3. Otway Basin<br />

Grange Burn, at 180 m elevation, near Hamilton in south-western Victoria is located close to<br />

the limits of marine transgression in the Otway Basin during Late Neogene time. The site<br />

provides a record of an Early Pliocene environment that is unique in Australia for two<br />

reasons. A regressive non-marine alluvium is preserved between a late Miocene-Early<br />

Pliocene marine silt and a K/AR-dated 4.46 Ma old basalt. Fossils preserved in the alluvium<br />

include the remains of plants and mammals animals whose NLRs are confined to coastal<br />

northeastern Queensland (Flannery et al. 1992, Macphail 1996b). Commonly occurring<br />

microfossils include Cyathea, Araucariaceae (Araucaria), Podocarpaceae (Dacrycarpus,<br />

Podocarpus-Prumnopitys), Casuarinaceae and Myrtaceae (including Eucalyptus). Less<br />

common or rare taxa are Dacrydium, Asteraceae, Eucalyptus spathulata-type, Nothofagus<br />

(Brassospora, Fuscospora, Lophozonia) spp. and Poaceae.<br />

Inferred climate<br />

The combined microfloral and palaeontological evidence point to a mosaic vegetation of<br />

Araucariaceae (dry) rainforest and open-canopy sclerophyll communities, which may have<br />

been confined to the stream banks. There is no evidence that Nothofagus spp. were part of<br />

these communities or present elsewhere in the district. Poaceae are suggested to have<br />

colonised drier (interfluve) areas but there is no evidence that grasslands per se were part of<br />

the regional dryland vegetation. Macphail (1996b) has proposed that climates were wet<br />

(humid) and weakly seasonal. Mean annual temperatures are likely to have been very warm<br />

(upper mesotherm).<br />

4. Murray Basin<br />

Marine transgressions during the Late Miocene and Early Pliocene resulted in the deposition<br />

of marginal marine facies in the south-west sector of the basin. Examples are the<br />

Bookpurnong Beds and basal Loxton-Parilla sands (Brown and Stephenson 1991).<br />

Microfloras from restricted marine and estuarine facies are dominated by Araucariaceae,<br />

Casuarinaceae and/or Myrtaceae, with relatively low and diminishing relative abundances of<br />

Nothofagus (Brassospora) spp. Microfloras from paralic facies where the marine influence<br />

was weak or absent are dominated by Nothofagus (Brassospora), Araucariaceae and<br />

Casuarinaceae, with lower abundances of Myrtaceae. These facies are difficult to correlate<br />

using plant microfossils and it is uncertain whether differences in pollen dominance are due to<br />

the Neves Effect, edaphic-forcing or evidence that Nothofagus-dominated rainforest was<br />

succeeded by drier, Araucariaceae-dominated, types of rainforest.<br />

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