OFR 151.pdf - CRC LEME

OFR 151.pdf - CRC LEME OFR 151.pdf - CRC LEME

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impression of floristic impoverishment is valid or merely due to cursory examination of the material. If the former then it is probable that conditions within the Bass Basin were as wet (perhumid) but cooler (upper microtherm) than in the Gippsland Basin. 2. North-west Coast Diverse microfloras are preserved in mudstones at Wilsons Creek in the lower Mersey Valley in northwestern Tasmania, and a rafted clast of estuarine mud deposited in offshore marls at Fossil Bluff at Wynyard (Macphail 1996a, M.K. Macphail unpubl. results). The Wilsons Creek microfloras lack Cyatheacidites annulatus but include two rare taxa (Mutisiaepollis patersonii, Tricolpites stylidioides) that first appear in the earliest Oligocene/latest Eocene Lemonthyme Creek and Wilmot Dam sections (Macphail and Hill 1994) and are likely to be no older than Early Oligocene. Dominance is shared between Lagarostrobos (24%), Podosporites-Prumnopitys (21%) and Nothofagus (Brassospora) spp. (15%), with lower relative abundances of Nothofagus (Nothofagus) sp. (9%), N. (Fuscospora) spp. (3%), Osmundaceae (2%), Araucaria (3%), Dacrycarpus (3%), Casuarinaceae (4%), Proteaceae (3%), Trimeniaceae (3%) and unidentified tricolporates (2%). Rare taxa include Sphagnum, Cyathea, Dicksonia, a possible pteridosperm (Alisporites simplis), Araucariaceae (Agathis/Wollemia), Podocarpaceae (Dacrydium, Microstrobos, Phyllocladus), Cyperaceae, Droseraceae, Ericales, Euphorbiaceae (Austrobuxus-type, Croton-type), Loranthaceae, Menyanthaceae, Myrtaceae and Sapotaceae. Foraminiferal evidence indicates that the Proteacidites tuberculatus Zone microflora from Fossil Bluff was deposited during the earliest Miocene Foram Zone N5. Nothofagus (Brassospora) spp. are abundant (34%). Frequent to common taxa are Cyathea (10%), Dicksoniaceae (10%), Osmundaceae (2%), Araucaria (9%), Dacrydium (5%), Lagarostrobos (3%), Podocarpus-Prumnopitys (7%), Nothofagus (Lophozonia) spp. (4%) and N. (Fuscospora) spp. (2%). Rare taxa include Lophosoria, Cupressaceae-Taxodiaceae, Podocarpaceae (Dacrycarpus, Halocarpus, Microcachrys, Microstrobos, Phyllocladus, Podosporites), Apocynaceae (Alyxia), Chenopodiaceae-Amarathaceae, Ascarina, Ericales, Euphorbiaceae (Amperea), Ilex, Myrtaceae (Austromyrtus-type, Eucalyptus), Proteaceae (Agastachys-type, Beauprea, Embothrium, Xylomelum occidentale-type), Quintinia, Sapindaceae (Dodonaea ericifolia-type, D. triquetra-type) and Winteraceae. The microfloras represent floristically complex evergreen rainforest lining coastal rivers and possible strandline communities (Fossil Bluff). Although the cryptogam and gymnosperm components are relatively diverse, the number of angiosperms is lower than in co-eval rainforest stands in mainland south-east Australia. Inferred climate Conditions are likely to have been cool (microtherm range) and uniformly wet to very wet (perhumid). 3. North-east Proteacidites tuberculatus Zone lignites at Pioneer preserve rich microfossil assemblages as well as macrofossils including tree trunks (Harris 1964b, 1965c, Hill and Macphail 1983). Dominant taxa are Nothofagus (Brassospora) spp. (49-64%) associated with lower numbers of Blechnaceae (2-3%), trilete types (1-3%), Dacrydium (2-4%), Nothofagus (Lophozonia) spp. (up to 5%), Nothofagus (Fuscospora) spp. (up to 4%), Proteaceae (1-3%) and unidentified tricolporate types (2-3%). Cyathea (6%), Quintinia (2%) and Trimeniaceae (5%) are occasionally frequent. Osmundaceae, Casuarinaceae, Cunoniaceae, Myrtaceae and Restionaceae are present in low but significant numbers (~1%). Rare taxa include: 265

Lophosoria, Dicksonia, Araucariaceae (Araucaria, Agathis/Wollemia), Podocarpaceae (Dacrycarpus, Halocarpus, Microstrobos, Phyllocladus), Anacardiaceae, Cupanieae, Elaeocarpaceae, Euphorbiaceae (Austrobuxus-type), Ilex, Loranthaceae, Myrtaceae (including Austromyrtus-type, Eucalyptus), Proteaceae (including Agastachys-type, Beauprea, Xylomelum occidentale-type), Sapotaceae and Winteraceae. The assemblages represent Nothofagus-gymnosperm temperate rainforest. A comparison of the macrofossil and microfossil data shows that the canopy dominants were species within the Nothofagus subgenus Lophozonia, not the subgenus Brassospora, as indicated by relative pollen abundance. The corollary is that Nothofagus (Brassospora) spp. are over-represented by pollen. Inferred climate Climates were upper microtherm and uniformly very wet (perhumid). 4. West Coast Lignites preserved in Tertiary palaeochannels near Zeehan and Balfour provide ‘snap shots’ of the Proteacidites tuberculatus Zone vegetation on the west coast (M.K. Macphail unpubl. results). Both microfloras represent floristically simple Nothofagus-gymnosperm evergreen rainforest. The possibly older Zeehan microflora is dominated by Nothofagus (Brassospora) spp., Proteaceae (especially Agastachys-type) and Casuarinaceae. Lagarostrobos, Quintinia, Trimeniaceae and a small Ilex grain are frequent. Rare taxa include Microstrobos, Gunnera, Lactoridaceae (now endemic to the Juan Fernandez Islands off Chile), Myrtaceae, Nothofagus (Lophozonia) and Strasburgeriaceae. Pollen dominance at the possibly younger Balfour site is shared between Nothofagus (Brassospora) spp. (30%), Dacrydium (10%), Lagarostrobos (15%), Podocarpus-Prumnopitys (11%), Casuarinaceae (6%) and Nothofagus (Fuscospora, Nothofagus) spp. (5%). Frequent types include Araucaria (2%), Podosporites (2%), Strasburgeriaceae (1%), Trimeniaceae (3%) and Ericales (2%). Rare taxa include Lophosoria, Cupressaceae, Gunnera, Polygalaceae, Quintinia and Winteraceae. Cupanieae are absent. Inferred climate Climates were cool (microtherm range) and uniformly wet (perhumid). Conditions do not appear to have been significantly cooler at Zeehan than at Balfour despite differences in elevation. 5. Central Plateau Sites preserving diverse Oligo-Miocene microfloras occur at Cethana, Lemonthyme Creek, Lea River, Link Road, Little Rapid River and Wilmot Dam in northwestern Tasmania (Macphail et al. 1993, Macphail and Hill 1994, M.K. Macphail unpubl. results) and at Monpeelyata, near Great Lake on the eastern margin of the Central Plateau (Macphail et al. 1991). In all cases the assemblages accumulated in palaeochannels incised into Jurassic or older rocks. The microfloras represent Nothofagus-gymnosperm rainforests, which were structurally less complex and floristically simpler than comparable rainforests growing in the south-east of the mainland. The earliest microfloras (Lemonthyme Creek and Wilmot Dam) are associated with probable earliest Oligocene glacial deposits (Lemonthyme Glaciation). K/Ar dates of overlying basalts confirm the sections are older than 26.7 Ma. Although assigned to the Proteacidites tuberculatus Zone, based on Cyatheacidites annulatus, it is probable that this fern 266

Lophosoria, Dicksonia, Araucariaceae (Araucaria, Agathis/Wollemia), Podocarpaceae<br />

(Dacrycarpus, Halocarpus, Microstrobos, Phyllocladus), Anacardiaceae, Cupanieae,<br />

Elaeocarpaceae, Euphorbiaceae (Austrobuxus-type), Ilex, Loranthaceae, Myrtaceae (including<br />

Austromyrtus-type, Eucalyptus), Proteaceae (including Agastachys-type, Beauprea,<br />

Xylomelum occidentale-type), Sapotaceae and Winteraceae. The assemblages represent<br />

Nothofagus-gymnosperm temperate rainforest.<br />

A comparison of the macrofossil and microfossil data shows that the canopy dominants were<br />

species within the Nothofagus subgenus Lophozonia, not the subgenus Brassospora, as<br />

indicated by relative pollen abundance. The corollary is that Nothofagus (Brassospora) spp.<br />

are over-represented by pollen.<br />

Inferred climate<br />

Climates were upper microtherm and uniformly very wet (perhumid).<br />

4. West Coast<br />

Lignites preserved in Tertiary palaeochannels near Zeehan and Balfour provide ‘snap shots’<br />

of the Proteacidites tuberculatus Zone vegetation on the west coast (M.K. Macphail unpubl.<br />

results). Both microfloras represent floristically simple Nothofagus-gymnosperm evergreen<br />

rainforest.<br />

The possibly older Zeehan microflora is dominated by Nothofagus (Brassospora) spp.,<br />

Proteaceae (especially Agastachys-type) and Casuarinaceae. Lagarostrobos, Quintinia,<br />

Trimeniaceae and a small Ilex grain are frequent. Rare taxa include Microstrobos, Gunnera,<br />

Lactoridaceae (now endemic to the Juan Fernandez Islands off Chile), Myrtaceae, Nothofagus<br />

(Lophozonia) and Strasburgeriaceae. Pollen dominance at the possibly younger Balfour site<br />

is shared between Nothofagus (Brassospora) spp. (30%), Dacrydium (10%), Lagarostrobos<br />

(15%), Podocarpus-Prumnopitys (11%), Casuarinaceae (6%) and Nothofagus (Fuscospora,<br />

Nothofagus) spp. (5%). Frequent types include Araucaria (2%), Podosporites (2%),<br />

Strasburgeriaceae (1%), Trimeniaceae (3%) and Ericales (2%). Rare taxa include<br />

Lophosoria, Cupressaceae, Gunnera, Polygalaceae, Quintinia and Winteraceae. Cupanieae<br />

are absent.<br />

Inferred climate<br />

Climates were cool (microtherm range) and uniformly wet (perhumid). Conditions do not<br />

appear to have been significantly cooler at Zeehan than at Balfour despite differences in<br />

elevation.<br />

5. Central Plateau<br />

Sites preserving diverse Oligo-Miocene microfloras occur at Cethana, Lemonthyme Creek,<br />

Lea River, Link Road, Little Rapid River and Wilmot Dam in northwestern Tasmania<br />

(Macphail et al. 1993, Macphail and Hill 1994, M.K. Macphail unpubl. results) and at<br />

Monpeelyata, near Great Lake on the eastern margin of the Central Plateau (Macphail et al.<br />

1991). In all cases the assemblages accumulated in palaeochannels incised into Jurassic or<br />

older rocks. The microfloras represent Nothofagus-gymnosperm rainforests, which were<br />

structurally less complex and floristically simpler than comparable rainforests growing in the<br />

south-east of the mainland.<br />

The earliest microfloras (Lemonthyme Creek and Wilmot Dam) are associated with probable<br />

earliest Oligocene glacial deposits (Lemonthyme Glaciation). K/Ar dates of overlying basalts<br />

confirm the sections are older than 26.7 Ma. Although assigned to the Proteacidites<br />

tuberculatus Zone, based on Cyatheacidites annulatus, it is probable that this fern<br />

266

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