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OFR 151.pdf - CRC LEME

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genera are Gymnostoma (Casuarinaceae) and Eucryphia (Eucryphiaceae); extinct genera<br />

include Banksieaephyllum (Proteaceae) (Hill 1991, Carpenter et al. 1994b, Scriven and Hill<br />

1995, Taylor and Hill 1996).<br />

The flora is an important one, not only because it includes taxa whose NLRs are widely<br />

separated in the South-west Pacific region, but also because of the diversity of morphological<br />

adaptations. For example the closest relatives of Gymnostoma (G. antiquum) and<br />

Banksieaephyllum (B. taylorii) occur in relatively dry areas in (tropical) New Guinea and<br />

(humid) southwestern Western Australia, respectively. The latter is the earliest known<br />

example of scleromorphy recorded in an Australia Tertiary angiosperm.<br />

Inferred climate<br />

Well-defined growth rings in the fossil gymnosperm wood point to tree growth being<br />

controlled by strong seasonal fluctuation in mean temperature since neither photoperiod nor<br />

precipitation are likely to have been limiting factors. Based on bioclimatic data from extant<br />

NLRs, Taylor et al. (1990) propose that mean annual temperatures were mesotherm (~14-<br />

20 0 C) but frost-free. To what degree the cool/wet conditions at Palaeolake Bungarby were<br />

due to elevation and orographic cloudiness reducing evapo-transpiration on the Southeastern<br />

Highlands of New South Wales is uncertain.<br />

1.1.7 Tasmania<br />

No known record.<br />

1.2 Microfloras<br />

Early and/or Late Paleocene marine sediments are widespread in the offshore Carnarvon,<br />

Canning and Bonaparte Basins but are seldom sampled (Apthorpe 1988). Dredge samples<br />

from the continental shelf off the central and northern New South Wales coast include some<br />

rocks that preserve Paleocene dinoflagellate floras (R. Helby pers. comm.) but information on<br />

the associated spore-pollen content is unavailable. Other Paleocene sequences in the same<br />

category occur off the western margin of Tasmania margin, and in north-west Tasmania (e.g.<br />

Sorell Basin) and south-west Western Australia (Kings Park Shale). For this reason what is<br />

currently known about the Paleocene flora and vegetation mostly comes from<br />

Lygistepollenites balmei Zone microfloras preserved in subcrop in southeastern Australia.<br />

1.2.1 North-West Australia<br />

No known record.<br />

1.2.2 North-East Australia<br />

1. Northern Tablelands of New South Wales<br />

Palaeosols and braided stream deposits underlie latest Paleocene [~54 Ma] basalts at<br />

elevations above 800 m on the Mt. Royal Range and sites near Tamworth on the Northeastern<br />

Tablelands (Morgan 1978, Martin et al. 1987).<br />

Microfloras recovered from Mt. Royal are dominated by araucarians (Araucaria), podocarps<br />

(Dacrydium, Lagarostrobos, Podocarpus-Prumnopitys) and ferns (Cyatheaceae,<br />

Gleicheniaceae). Angiosperm pollen are relatively rare, with the most common taxon being<br />

small, nondescript Proteaceae. Nothofagus is absent. The Tamworth assemblages are<br />

dominated by Podocarpaceae (Dacrycarpus, Lagarostrobos, Podocarpus-Prumnopitys), but<br />

204

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