OFR 151.pdf - CRC LEME

OFR 151.pdf - CRC LEME OFR 151.pdf - CRC LEME

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2. TIME SLICE K-2 Age Range: Aptian – Albian [115- 97.5 Ma] Zones: Cyclosporites hughesii to Phimopollenites pannosus Zones Odontochitina operculata to lower Xenascus asperatus Zones 2.1 Macrofloras Basins that preserve macrofossil records for the Berriasian-Barremian usually preserve Aptian-Albian macrofloras as well. However the only comprehensive records come from basins located along the present-day southern margin, then at high palaeolatitude (70-80 0 S) along the palaeo-western margin (see Figure 5). 2.1.1 North-West Australia No known record. 2.1.2 North-East Australia 1. Marlborough Basin A sparse macroflora preserved in the Marlborough Basin, southern Queensland, indicates that forests or woodlands on the northernmost of the four islands making up Australia during the Aptian included Araucariaceae, Ginkgoites, pteridosperms/Pentoxylales (Taeniopteris), sphenopsids and ferns. Dettmann and Clifford (2000a) have recorded propagules resembling gemmae produced by modern liverworts (Marchantiales) in latest Albian sediments from the south-east of the Eromanga Basin. Inferred climate Since the Marlborough Basin was located at a palaeolatitude of about 75 0 S, it is reasonable to assume that conditions were humid and temperatures were below freezing (lower microtherm) during winter. 2.1.3 Central Australia 1. Eromanga and Surat Basins Palaeogeographic reconstructions indicate the depocentre of the Eromanga Basin was located at about 60 0 S. The Euromanga and Surat Basins to the east were part of an extensive epicontinental seaway stretching from the palaeo-northern margin at about ~50 0 S southwards to about palaeolatitude 80 0 S. Silicified wood with narrow growth rings is preserved in carbonate concretions in the Early Albian, upper Bulldog Shale and Toolebuc Formation and a cycad is preserved in the Middle-Late Albian Allaru Mudstone in the Eromanga Basin but other macrofossils from this and the Surat Basin await identification (references in Dettmann et al. 1992). 175

Inferred climate The stems are believed to represent podocarp-dominated forests or woodlands (Austral Conifer Forest) growing under wet (humid) but strongly seasonal and cool-cold (microtherm range) conditions on uplands surrounding the Eromanga Basin (Francis and Frakes 1993). 2.1.4 South-West Australia 1. Carnarvon and Perth Basins Fossil macrofloras consisting of permineralised wood are preserved in the Early Aptian Muderong Shale and Middle Aptian-Albian Windalia Radiolarite in the Carnarvon Basin, and Early Aptian Dandaragan Sandstone in the Perth Basin (McLoughlin 1996). Inferred climate McLoughlin (ibid) suggests that araucarians and podocarps are likely to have dominated tree communities, implying conditions were humid. Based on the middle palaeolatitude location (~55 0 S) on the palaeo-northern margin, mean annual temperatures may have been relatively warm (lower mesotherm) and seasonal contrasts less marked than in inland regions. 2.1.5 Central southern Australia No known record. 2.1.6 South-East Australia Aptian to Early Albian (Zone C) and Early to Late Albian (Zone D) macrofloras are widely preserved across the Otway and Gippsland Basins, which were located on the palaeo-western margin between latitudes 70-80 0 S (Douglas 1969, 1973a, 1986, 1994). The most important of these floras is a Zone C macroflora preserved in the Koonwarra Beds, onshore Gippsland Basin. The Zone C/D boundary approximates to the Aptian/Albian boundary. 1. Aptian to Early Albian The Koonwarra Beds preserve a particularly diverse record of the earliest Aptian plants and animals living in and around a large (>2 km diameter) freshwater lake (Drinnan and Chambers 1986, Rich et al. 1988). The flora includes most of the major plant taxa forming Austral Conifer Forest elsewhere in southern Australia during the early Cretaceous. These plants ranged in size from bryophytes to probably tall gymnosperms. Angiospermids are represented by a flower (Taylor and Hickey 1990) and fossil pollen. Important taxa include Araucariaceae, Podocarpaceae, Ginkgoites, pteridosperms/Pentoxylales (Taeniopteris), ferns and fern allies (Adiantites, Gleichenites), sphenopsids (Equisetum, Phyllotheca, Sphenopteris), isoetaleans (including Isoetes), lycopods (Lycopodium), mosses (including Sphagnales) and liverworts (Jungermanniales, Anthocerotae). The woody flora included both deciduous and evergreen taxa. Many of the sphenopsids, ferns and fern allies closely resemble their nearest living relatives, e.g. Equisetum, Gleichenia and Lycopodium. The bryophyte flora included ground and epiphytic species. The remains of small ‘polar’ dinosaurs and birds are preserved in the Koonwarra Beds and correlative deposits elsewhere in southern Victoria. 176

Inferred climate<br />

The stems are believed to represent podocarp-dominated forests or woodlands (Austral<br />

Conifer Forest) growing under wet (humid) but strongly seasonal and cool-cold (microtherm<br />

range) conditions on uplands surrounding the Eromanga Basin (Francis and Frakes 1993).<br />

2.1.4 South-West Australia<br />

1. Carnarvon and Perth Basins<br />

Fossil macrofloras consisting of permineralised wood are preserved in the Early Aptian<br />

Muderong Shale and Middle Aptian-Albian Windalia Radiolarite in the Carnarvon Basin, and<br />

Early Aptian Dandaragan Sandstone in the Perth Basin (McLoughlin 1996).<br />

Inferred climate<br />

McLoughlin (ibid) suggests that araucarians and podocarps are likely to have dominated tree<br />

communities, implying conditions were humid. Based on the middle palaeolatitude location<br />

(~55 0 S) on the palaeo-northern margin, mean annual temperatures may have been relatively<br />

warm (lower mesotherm) and seasonal contrasts less marked than in inland regions.<br />

2.1.5 Central southern Australia<br />

No known record.<br />

2.1.6 South-East Australia<br />

Aptian to Early Albian (Zone C) and Early to Late Albian (Zone D) macrofloras are widely<br />

preserved across the Otway and Gippsland Basins, which were located on the palaeo-western<br />

margin between latitudes 70-80 0 S (Douglas 1969, 1973a, 1986, 1994). The most important of<br />

these floras is a Zone C macroflora preserved in the Koonwarra Beds, onshore Gippsland<br />

Basin. The Zone C/D boundary approximates to the Aptian/Albian boundary.<br />

1. Aptian to Early Albian<br />

The Koonwarra Beds preserve a particularly diverse record of the earliest Aptian plants and<br />

animals living in and around a large (>2 km diameter) freshwater lake (Drinnan and<br />

Chambers 1986, Rich et al. 1988).<br />

The flora includes most of the major plant taxa forming Austral Conifer Forest elsewhere in<br />

southern Australia during the early Cretaceous. These plants ranged in size from bryophytes<br />

to probably tall gymnosperms. Angiospermids are represented by a flower (Taylor and<br />

Hickey 1990) and fossil pollen. Important taxa include Araucariaceae, Podocarpaceae,<br />

Ginkgoites, pteridosperms/Pentoxylales (Taeniopteris), ferns and fern allies (Adiantites,<br />

Gleichenites), sphenopsids (Equisetum, Phyllotheca, Sphenopteris), isoetaleans (including<br />

Isoetes), lycopods (Lycopodium), mosses (including Sphagnales) and liverworts<br />

(Jungermanniales, Anthocerotae).<br />

The woody flora included both deciduous and evergreen taxa. Many of the sphenopsids, ferns<br />

and fern allies closely resemble their nearest living relatives, e.g. Equisetum, Gleichenia and<br />

Lycopodium. The bryophyte flora included ground and epiphytic species. The remains of<br />

small ‘polar’ dinosaurs and birds are preserved in the Koonwarra Beds and correlative<br />

deposits elsewhere in southern Victoria.<br />

176

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