OFR 151.pdf - CRC LEME

OFR 151.pdf - CRC LEME OFR 151.pdf - CRC LEME

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fossil taxa that are morphologically indistinguishable from extant species, almost certainly represent extinct ecotypes (Macphail et al. 1993, Jordan 1997a). 7.8.3 Palaeoclimates Australian marine records for the Late Neogene are mostly confined to the north, mainly the Coral Sea (Feary et al. 1991) and North West Shelf (Quilty 1994). These imply that Pliocene climates were gradually cooling, although still warm and generally stable. Palaeobotanical evidence for Late Neogene climates in Australia are reviewed by Kershaw et al. (1994) and Macphail (1997b). As for earlier periods within the Tertiary, the age control is of variable quality, and reconstruction of Late Neogene palaeotemperatures depends heavily on the assumption that Late Neogene species of Araucaria and Agathis had much the same temperature requirements as their NLRs in Australia. On present indications, patterns of wind and oceanic circulation became comparable to those of the present-day during the late Neogene whilst geographic variation in the relative abundance and time of extinction of the ‘Tertiary rainforest’ component provides a reliable indication of the direction and changing strength of climatic gradients across the continent (Macphail 1997b). Some Late Neogene vegetation types have no surviving analogue in Australia, and the same may be true of the associated microclimates. Palaeo-northern Australia Sparse records from northwestern Australia (Macphail 1997b) indicate that mean annual rainfall decreased from 600-1500 mm to below 600 mm and became strongly seasonal (aridsemiarid) during the Late Miocene, and this drying trend was maintained throughout the Pleistocene. Lowland climates in coastal northeastern Australia were seasonally too dry and/or warm to support Nothofagus (Brassospora) spp. during the Late Miocene although cooler (lower mesotherm) conditions allowed small populations to survive at high elevations on the Atherton Tableland into Pliocene time. On the coast, summer rainfall and mean annual temperatures were adequate (upper mesotherm, perhumid) to support a mosaic of Araucariaceae dry rainforest and angiosperm-dominated sclerophyll communities during the Late Miocene, but an increasing representation of sclerophyll taxa and hints that rainfall became increasingly seasonal (possibly monsoonal) during the Pliocene. The combined data imply that a very strong west to east rainfall gradient – from 1300 mm pa – was in existence across northern Australia by the Late Pliocene. Palaeo-central Australia Deep weathering during the Neogene has destroyed all organic remains except in highly fortuitous contexts. The limited data available (including macrofaunas) point to the continuing existence of permanent or semi-permanent freshwater ponds. Although vegetation is likely to have been open sclerophyll woodland or shrubland, some evergreen rainforest species survived in gallery communities lining possibly ephemeral rivers draining into the Eyre Basin. The regional climate appears to have been humid to sub-humid (below 900-1000 mm pa) and strongly seasonal. Palaeo-southern Australia A comparison of microfloras from southwestern and southeastern Australia confirms that rainfall increased markedly along west to east gradients during the Late Neogene; the reliability and amount of rainfall received during summer months almost certainly increased along the same longitudinal gradient as well as upslope. Similarly, a comparison of sites in northern New South Wales and Tasmania confirms the decrease in mean annual temperature with increasing latitude. Similar gradients are recorded upslope in the Northern Tablelands and Southeastern Highlands of New South Wales and were presumably also present in 107

mountainous regions of Tasmania. Data from individual sites that preserve vegetation histories covering all or part of Late Miocene to Late Pliocene time flesh-out the palaeoclimatic inferences. A particularly detailed record of climatic change in south-west Western Australia during the mid Pliocene warm period is preserved in a probable meteor impact crater site at Yallalie north of Perth (Dodson and Ramrath in press, Dodson and Macphail in press). 1. Rainfall The survival of Araucaria, Agathis, Dacrycarpus, Dacrydium and Nothofagus (Brassospora) spp. in south-west Western Australia (Yallalie), southwestern New South Wales (Lachlan Valley), southwestern Victoria (Grange Burn), southeastern Victoria (Hapuku-1), and western Tasmania (Linda Valley) is compelling evidence that conditions during the mid Pliocene were effectively wetter than at present although at Yallalie and possibly Grange Burn groundwater discharge may have compensated for any moisture deficit during summer. Transient upsurges in the relative abundance of semiarid plants and halophytic diatoms at Yallalie demonstrate that warm (upper mesotherm) and seasonally wet conditions within and beyond the crater were interrupted by three distinct episodes of aridification at about 2.90 Ma, 2.59 Ma and 2.56 Ma. In contrast, if stands of Araucariaceae dry rainforest at Grange Burn were wholly supported by rainwater, then mean annual rainfall will have been between 1300-2000 mm versus 680 mm at present. Trends in relative pollen abundance in the Murray and Gippsland Basins imply that conditions in south-eastern Australia remained relatively wet (humid) during the Early Pliocene, and rainfall became weakly to strongly seasonal during the Late Pliocene. The apparent increase in the mean annual precipitation on the Southeastern Highlands of New South Wales (Lachlan Valley, Lake George) is likely to be local, due to orographic cloudiness reducing evapo-transpiration losses. 2. Temperature The survival or re-establishment of Araucariaceae at Grange Burn before 4.46 Ma and Yallalie between 2.5-3.6 Ma indicates that mean temperatures in south-west and central southern Australia during the Early to Middle Pliocene were within the upper mesotherm range. If the climatic envelope for the southern-most living species of Agathis (A. bidwillii) is used as a guide, then the mean minimum temperature of the coldest month will have been about 5-10 0 C and mean maximum temperature of the hottest month will have been about 28- 32 0 C. Slightly lower mean values are obtained if the climatic data for living Araucaria are used. Because of site-specific factors, palaeotemperatures in southeastern Australia are more difficult to estimate in absolute terms. However geographic and temporal changes in the relative abundance of Araucariaceae indicate: (1) Mean values (lower mesotherm) were lower than in southwestern and central southern Australia. Cool-cold (microtherm) conditions occurred at high elevations on the Southeastern Highland and Tasmania. (2) Mean temperatures during the Early Pliocene were generally warmer (mesotherm range) than in the Late Pliocene (microtherm range). 3. Late Neogene climatic instability Oxygen isotope data indicate that global climates became increasingly variable (unstable) during the Late Neogene. Two sites that appear to link the aridification of Australia with increasingly variable climates are diatomites preserved in Palaeolake Bunyan on the Southeastern Highlands of New South Wales, and Yallalie (ibid). Taylor et al. (1990) propose that cyclic bedding of the Bunyan diatomite is evidence for climatic cycles with a periodicity less than or equal to 8000 years during the Late Miocene in southeastern Australia. The arid vegetation response seen at Yallalie is similar to that seen in southeastern Australia 108

mountainous regions of Tasmania. Data from individual sites that preserve vegetation<br />

histories covering all or part of Late Miocene to Late Pliocene time flesh-out the<br />

palaeoclimatic inferences. A particularly detailed record of climatic change in south-west<br />

Western Australia during the mid Pliocene warm period is preserved in a probable meteor<br />

impact crater site at Yallalie north of Perth (Dodson and Ramrath in press, Dodson and<br />

Macphail in press).<br />

1. Rainfall<br />

The survival of Araucaria, Agathis, Dacrycarpus, Dacrydium and Nothofagus (Brassospora)<br />

spp. in south-west Western Australia (Yallalie), southwestern New South Wales (Lachlan<br />

Valley), southwestern Victoria (Grange Burn), southeastern Victoria (Hapuku-1), and western<br />

Tasmania (Linda Valley) is compelling evidence that conditions during the mid Pliocene were<br />

effectively wetter than at present although at Yallalie and possibly Grange Burn groundwater<br />

discharge may have compensated for any moisture deficit during summer. Transient upsurges<br />

in the relative abundance of semiarid plants and halophytic diatoms at Yallalie demonstrate<br />

that warm (upper mesotherm) and seasonally wet conditions within and beyond the crater<br />

were interrupted by three distinct episodes of aridification at about 2.90 Ma, 2.59 Ma and 2.56<br />

Ma. In contrast, if stands of Araucariaceae dry rainforest at Grange Burn were wholly<br />

supported by rainwater, then mean annual rainfall will have been between 1300-2000 mm<br />

versus 680 mm at present. Trends in relative pollen abundance in the Murray and Gippsland<br />

Basins imply that conditions in south-eastern Australia remained relatively wet (humid)<br />

during the Early Pliocene, and rainfall became weakly to strongly seasonal during the Late<br />

Pliocene. The apparent increase in the mean annual precipitation on the Southeastern<br />

Highlands of New South Wales (Lachlan Valley, Lake George) is likely to be local, due to<br />

orographic cloudiness reducing evapo-transpiration losses.<br />

2. Temperature<br />

The survival or re-establishment of Araucariaceae at Grange Burn before 4.46 Ma and<br />

Yallalie between 2.5-3.6 Ma indicates that mean temperatures in south-west and central<br />

southern Australia during the Early to Middle Pliocene were within the upper mesotherm<br />

range. If the climatic envelope for the southern-most living species of Agathis (A. bidwillii) is<br />

used as a guide, then the mean minimum temperature of the coldest month will have been<br />

about 5-10 0 C and mean maximum temperature of the hottest month will have been about 28-<br />

32 0 C. Slightly lower mean values are obtained if the climatic data for living Araucaria are<br />

used.<br />

Because of site-specific factors, palaeotemperatures in southeastern Australia are more<br />

difficult to estimate in absolute terms. However geographic and temporal changes in the<br />

relative abundance of Araucariaceae indicate: (1) Mean values (lower mesotherm) were lower<br />

than in southwestern and central southern Australia. Cool-cold (microtherm) conditions<br />

occurred at high elevations on the Southeastern Highland and Tasmania. (2) Mean<br />

temperatures during the Early Pliocene were generally warmer (mesotherm range) than in the<br />

Late Pliocene (microtherm range).<br />

3. Late Neogene climatic instability<br />

Oxygen isotope data indicate that global climates became increasingly variable (unstable)<br />

during the Late Neogene. Two sites that appear to link the aridification of Australia with<br />

increasingly variable climates are diatomites preserved in Palaeolake Bunyan on the<br />

Southeastern Highlands of New South Wales, and Yallalie (ibid). Taylor et al. (1990)<br />

propose that cyclic bedding of the Bunyan diatomite is evidence for climatic cycles with a<br />

periodicity less than or equal to 8000 years during the Late Miocene in southeastern Australia.<br />

The arid vegetation response seen at Yallalie is similar to that seen in southeastern Australia<br />

108

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