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OFR 151.pdf - CRC LEME

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Similarly it is difficult to summarise bioclimates within most of the selected geographic<br />

regions due to strengthening temperature and rainfall gradients between coastal, inland and<br />

upland districts. Nonetheless changes in miospore dominance and the palaeodistribution of<br />

some cryptogams provide unequivocal evidence of the impact of cooling followed by<br />

temporary warming during the Oligocene-Early Miocene. Examples include the marked<br />

impoverishment of Nothofagus forest in the onshore Gippsland Basin during the Eocene-<br />

Oligocene transition (Upper Nothofagidites asperus Zone) and the re-establishment of<br />

Lophosoria in southeastern Australia during the Early Oligocene (Macphail et al. 1994).<br />

Fossil spores of Lophosoria (Cyatheacidites annulatus) indicate this large ground fern, which<br />

is now restricted to cool-cold climates in South America, reached Tasmania via trans-oceanic<br />

dispersal during or slightly before the Lemonthyme Glaciation (Macphail and Hill 1994). The<br />

fern then spread rapidly northwards throughout southeastern Australia during the Early<br />

Oligocene (Proteacidites tuberculatus Zone time) but is not recorded in southern Queensland<br />

until the Early Miocene (Dettmann, 1986b), due to relatively warm conditions at lower<br />

latitudes. Increasingly dry, seasonal climates explain the failure of Lophosoria to extend into<br />

central and north-west Australia.<br />

Palaeo-northern Australia<br />

Sparse pollen evidence from the Argo abyssal plain off Port Hedland hints that continental<br />

climates in north-west Western Australia were strongly seasonal (upper mesothermmegatherm,<br />

semiarid-subhumid) although Oligocene to Early Miocene microclimates in the<br />

Pilbara region were sufficiently humid during summer months to allow Nothofagus<br />

(Brassospora) spp. and podocarps to survive along palaeochannels on the Hamersley Ranges<br />

(M.K. Macphail unpubl. data). Conditions in inland northeastern Australia appear to have<br />

been warm to hot (upper mesotherm) and seasonally very wet (perhumid) based on faunal<br />

remains and leaf impressions of subtropical-tropical rainforest species preserved in karst<br />

terrain at Riversleigh, northwestern Queensland (R.S. Hill pers. comm.).<br />

Isotopic evidence from the Coral Sea indicates SSTs in the region increased from between<br />

9.5-13.0 0 C (microtherm range) in the earliest Early Oligocene to 14.5-19.5 0 C (mesotherm<br />

range) in the Early Oligocene and reached a maximum of 20.5 0 C (upper mesotherm) in the<br />

Middle Miocene (Feary et al. 1991). Floristically complex Nothofagus communities<br />

remained the dominant vegetation type on the western slopes of the Leichhardt Range inland<br />

of Mackay, into Early-Middle Miocene (Canthiumidites bellus Zone Equivalent) time<br />

(Beeston 1994). This rainforest included taxa with cool temperate NLRs, e.g. Dacrydium,<br />

Lagarostrobos, Microcachrys and Nothofagus (Lophozonia), as well as taxa with warm<br />

temperate to subtropical-tropical NLRs, e.g. Anacolosa, Archidendron-type, Cupanieae and<br />

Ilex. How representative this flora and vegetation were of areas to the west in central<br />

Queensland is unclear. Further to the south, uniformly wet and relatively cool (lower<br />

mesotherm) climates allowed Nothofagus (Brassospora) spp. to become prominent during the<br />

Late Oligocene in coastal southern Queensland. Conditions became increasingly warm<br />

(upper mesotherm) and/or seasonally dry, resulting in the replacement of Nothofagus<br />

communities by other rainforest types during the Early to Middle Miocene. The presence of<br />

Nypa in Foram Zones N5-N7 marine sediments but not in Foram Zones N8-N9 marine<br />

sediments in the Capricorn Basin implies maximum warmth in southern Queensland occurred<br />

during the Early Miocene.<br />

The combined data demonstrate that precipitation gradients across northern Australia were<br />

parallel in direction but not as strong as those of the present-day. Because warm water flow<br />

through the Indonesian archipelago was not severely constricted until the Early Pliocene<br />

(Srinivasan and Sinha 1998), it is premature to explain these gradients in terms of modern<br />

ocean current patterns or monsoonal climates associated with uplift of the Tibetan Plateau (cf.<br />

Ramstein et al. 1997).<br />

101

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