Baden-Württemberg - Lichens of Wales

larger, thallus not radiate- rosetted with enlarged
marginal areoles, rust- brown to ochre)
8* Thallus not rust-red to rust brown 9
9 Thallus or ap. K+ red (crystals), C-. Norstictic
Acid 10
9* Thallus and ap. K-, C- or C+/KC+ red . 11
10 Thallus light gray-brown to light red-brown,
areolate to almost squamulose, areoles for the
most part taken up by ap., rounded to angular, -1
mm. Ap. K+ red, single, rarely to 2-5, flat or
concave, light margined, -4.0 mm. Sp. 4-6 x 1.5-
2 µm. * A. paupera
10* Thallus yellowish-brown, light brown, beige,
rarely red-brown or rust colored or greenish-
yellow, squamulose, K+ red or K-. Squamules
scattered or in small groups, rounded to irregular
and crenate, often with uneven upper surface,
mostly 0.5-2 mm, only a small part taken up by
ap. Ap. mostly 3-7, brown-black, punctiform,
rarely with disk opened out (mostly -0.8 mm).
Sp. 3-5 x 1-1.5(2) µm. * A. smaragdula
11 Thallus KC+ red. Gyrophoric acid 12
11* Thallus KC- 16
12 Thallus pruinose . 13
12* Thallus not pruinose . 14
13 Ap. 0.3-1.8 mm, disk brown to black-brown, very
uneven to umbilicate. With enlarged marginal
areoles, therefore appearing lobed. Areoles 0.4-
2.2 mm, convex, brown, often pruinose. Sp. 3-5
x 1.5-2 µm. *. On calcareous or neutral/basic
silicate rock - mieur; MRh
A. bullata Anzi
13* Ap. -0.4 mm, often only punctiform, hollowed
out, without recognizable margin, disk not
conspicuously uneven. Thallus squamulose to
areolate, dark brown to black- brown, squamules
05-3 mm, mostly in a commonly pruinose,
brown, very variable in size, thallus. Marginal
areoles often sterile, enlarged or not, sometimes
appearing lobed. Sp. 3-5 x 1.5-2 µm
* . A. umbilicata auct.
(In contrast to occasionally pruinose forms of A.
fuscata ap. smaller. Species concept unclear at
this time. The forms found in the region
probably not belonging to A. umbilicata Bagl.)
14 Ap. disk commonly smooth or very slightly
rough, not wrinkled, not umbilicate, ap. -1 mm,
hollowed out to level with the thallus upper
surface, mostly angular or irregular, often darker
than the thallus, red-brown to black-brown.
Thallus squamulose to (in the interior) areolate,
light to dark brown, or red-brown, usually ±
shiny, concave, flat or uneven, 0.5-1.5(3) mm.
Sp. 4-6 x 1-1.5 µm. Hym. 70-120 µm.
* . A. fuscata
14* Ap. disk wrinkled uneven or with umbilicus.
Hym. 80-100 µm. Very rare species 15
15 Ap. -0.3 (0.8) mm, mostly to 3-6, dark brown,
clearly margined, ± circular, hollowed out, disk
often with an umbilicus. squamules pale yellow-
brown, from flat, close standing to scattered, 0.4-
1.4 mm wide, ± pale underside squamules.
* ! A. gallica
15* Ap. -1.4 mm, dark red-brown, finally projecting
to sitting up, with erect thalloid margin and
wrinkled to often umbilicate disk, often close
standing. Thallus squamulose, squamules often
appearing lobed, shiny, light- to dark brown,
convex to uneven, 0.5-3 mm. Sp. 4-6 x 1.5-2
µm. * A. peliscypha
16 Sp. 9-13 x 5-6 µm, only to ca. 16-64 per ascus.
Areoles olive- to dark brown, undersides pale,
usually scattered and rounded, ± convex, 0.5-1
(1.5) mm. Ap. single, -0.6(1) mm, round, blackbrown
. A. oligospora
16* Sp. clearly smaller, to more than 64 per ascus . 17
17 Thallus pruinose 18
17* Thallus not pruinose 19
18 Thallus ± thickly pruinose, under the pruinosity
light brown to light red-brown, squamulose
areolate, squamules flat to convex, rounded to
angular, 0l5-2mm. Ap. ± hollowed, commonly
without projecting thalloid margin, usually blackbrown,
0.2-0.5 mm. Sp. 4-7 x 1.5-2 µm. Paraph.
ca. 1.5 µm thick. Hym. 90-130 µm
.A. versicolor
18* Thallus weakly to thickly pruinose, pruinosity
mostly truly variable in each thallus and only
occurring in the case of lime influence. Ap. often
with raised thalloid margin
. ↑ A. nitrophila (31)
19 Ap. -1.5(2) mm, red-brown to brown-black,
definite (usually raised) margin, to 1 (2-3) per
squamule, finally often clearly projecting and ±
sessile, rounded to distorted. squamules scattered
to running together, moderate to dark brown,
rounded, flat to convex. Hym. low, -75 µm high,
paraphyses -3.5 µm (above -5 µm) thick. sp. 3-6
x 1.5-2.5 µm . A. badiofusca
19* Ap. rarely over 0.8 mm. Hym higher. Paraph. -
2µm 20
20 Ap. commonly with warty rough disk, red- brown
thin margined, -0.8(1.4)mm. Hym. (120)140-
170 µm. Squamules 1-5 mm, yellow-brown,
light- to dark-brown, dull, usually uneven,
underside ± pale, usually scattered to in small
groups, often appearing lobed. Sp. 3.5-5.5 x 1.5-
2 µm A. scabrida
20* Ap. disk usually not warty-rough . 21
21 Thallus of single, ± rounded, -2 mm large,
weakly convex areoles or flowing together warty
areolate, yellow-brown, light brown, beige, often
with uneven upper surface, usually 0.5-2 mm,
underside ± blackish. Hym. (60)125-140(250)
µm. Ap. usually to 3-7, punctiform or rarely with
opened up disk (up to 0.8 mm), with or without
margin. Paraph. 1-15 µm thick. Sp. 2.5-5 x 1-
1.5(2) µm. Norstictic acid A. smaragdula
69