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Baden-Württemberg - Lichens of Wales

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Psoretum – s’mieur-med – v.rare (1); Mn<br />

(Taubertal), FrJu, Do, Av<br />

Catapyrenium waltheri (Krempelh.) Körber<br />

(Dermatocarpon w. (Krempelh.) Blomb. &<br />

Forss.)<br />

In alpine sites on naked soil and raw humus over<br />

calcareous subsoil – arct-alp – v.rare; Al;<br />

scattered in the Alps<br />

LIT: BREUSS 1990<br />

Catillaria Massal.<br />

(Keys include Catinaria, Cliostomum, Megalaria,<br />

Micarea p.p.)<br />

Introduction<br />

In the broad sense one places in Catillaria<br />

crustose lichens with two-celled colorless spores<br />

and various colored apothecia without thalloid<br />

margins. Catinaria, Cliostomum, Megalospora<br />

and Micarea were separated from this unnatural<br />

group genus according to various characteristics<br />

in the construction <strong>of</strong> the apothecium. Catillaria<br />

in the narrow sense includes species with<br />

generally thin, gray to nearly indistinguishable<br />

thalli living in the interior <strong>of</strong> the substrate,<br />

usually black to brown-black apothecia with<br />

lecideine proper margin, paraphyses unbranched<br />

to singly branched in the upper part with capitate<br />

thickened, provided with a brown apex capped<br />

ends and two-celled elongate spores.<br />

C. atomarioides and C. chalybeia occur on<br />

(mostly mineral rich) silicate rock; C.<br />

lenticularia lives on calcareous rocks and C.<br />

nigroclavata on bark. C. chalybeia frequently<br />

invades man made substrates. The indigenous<br />

rock dwellers are distributed in the south and<br />

central Europe and penetrate above all in the<br />

west widely into the boreal zone. C.<br />

nigroclavata occurs up into central<br />

Fennoscandia.<br />

C. detractula, C. erysiboides, C. minuta and C.<br />

picila do not belong to Catillaria s.str. The latter<br />

two live on shaded limestone on vertical and<br />

overhanging surfaces on humid habitats. C.<br />

detractula is found on well lighted, warm sites on<br />

limestone, C. erysiboides on tough old wood, e.g.<br />

deciduous tree stumps. C. erysiboides is<br />

distributed in the boreal and summer green<br />

deciduous region. The area <strong>of</strong> C. alba, C.<br />

anomaloides, C. detractula, C. picila and C.<br />

minuta stretches over south and (predominantly<br />

to the south) central Europe; finally an<br />

occurrence (Gotland) in south Scandinavia.<br />

Genus Characteristics and Determination<br />

Thallus crustose, clearly developed to living in<br />

the substrate, whitish, gray, brown, blackish, with<br />

various coccoid and filamentous green algae.<br />

Ap. variously colored (in Catillaria i.e.s. black to<br />

dark brown), without thalloid margin, with or<br />

rarely without definite proper margin. Exc.<br />

usually clearly developed. Paraphyses simple to<br />

sparsely branched, capitate above, with brown<br />

cap (Catillaria i.e.s.) or colorless to pigmented<br />

above. Hyp. colorless to brown-black. Asci<br />

almost cylindric to clavate, in the case <strong>of</strong><br />

Catillaria i.e.s. with uniformly I+ (K/I+) blue<br />

tholus and exterior with thin I+ (K/I+) blue<br />

gelatin layer, in the case <strong>of</strong> Catillaria i.w.s. to the<br />

part Biatora- and Bacidia- type. Sp. 2-celled.<br />

Ch-.<br />

1 On garden soil, humus, plant detritus, moss . 2<br />

1* On rock, bark, wood, living on conifer needles . 6<br />

2 On naked mineralized, sandy-loam to loamy<br />

soils. Ap. convex to hemispherical, marginless,<br />

black, -0.6 mm. Epihym. and hyp. dark purplebrown,<br />

hym. purple. Exc. dark purple, soon<br />

reduced. Sp. 1-2 celled, 9-15 x 3-4(4.5) µm.<br />

Paraph. branched in the upper part. Thallus very<br />

thin flowing, scattered or warty areolate, blue<br />

greenish. Saxony .<br />

Micarea melaenida (Nyl.) Coppins<br />

2* On mosses, plant detritus, raw humus, humus<br />

soils 3<br />

3 Hyp. light, mostly colorless. Ap. whitish,<br />

yellowish, rose, brown, discolored gray 4<br />

3* Hyp. red-brown to black-brown 5<br />

4 Sp. 10-17 x 4-5 µm, with definite epispore, 2- to<br />

rarely 4-celled. Ap. brown-rose, orange, roseyellowish,<br />

yellowish, beige, convex from the<br />

first, -0.9(1.1) mm, sometimes coalescing, margin<br />

rapidly disappearing. Epihym., hym., hyp. and<br />

exc. colorless to slightly yellowish, exc. strongly<br />

developed, paraplectenchymatous. Paraph.<br />

scarcely branched, strongly cemented, 3-4 µm.<br />

Thallus greenish-gray, thickly fine granular to<br />

erupting sorediate, the sorediate region greenish,<br />

soredia 30-70 µm. Zeorine . .<br />

Mycobilimbia sphaeroides<br />

4* Sp. smaller. Ap. convex. Paraph. branched and<br />

reticulate. Exc. mostly strongly reduced, paraph.<br />

hyphae like ↑ Micarea<br />

145

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