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Baden-Württemberg - Lichens of Wales

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quinone containing yellow to red colored part <strong>of</strong><br />

the Caloplaca species react blue-red with KOH.<br />

The genus in certain groups still<br />

taxonomically unclear is represented in central<br />

Europe by perhaps 50, in Germany with ca. 75<br />

species, <strong>of</strong> which however not a few are<br />

insufficiently known. The individual species<br />

reside on truly variable substrate, which mostly<br />

are however dependent on basic, subneutral or<br />

weakly acid conditions. Many are found on<br />

distinctly nutrient rich habitats, such as the<br />

calciphytic rock lichens C. citrina, C. decipiens,<br />

C. coronata, and C. ruderum. In addition C.<br />

alociza, C. arnoldii, c. aurantica, C. chalybaea,<br />

C. cirrochroa, c. dolomitica, C. flavescens, C.<br />

granulosa, C. holocarpa, C. lactea, C. ochracea,<br />

C. polycarpa, C. saxicola, C. variabilis and<br />

identified from the Allgäu C. nubigena and C.<br />

percrocata reside on calcareous rock, commonly<br />

in well lighted habitats. C. proteus is found<br />

predominantly on dolomite. The species C.<br />

teicholyta and C. flavovirescens are typically<br />

limited to calcareous rock. Such substrates are<br />

scarce in the region, however tomb stones, wall<br />

crowns and similar substrates have been used.<br />

On them, new habitats have been found by these<br />

and ecologically related species. C. teicholyta<br />

and C. crenulatella occur almost only on this<br />

kind <strong>of</strong> anthropogenic habitats.<br />

A greater part <strong>of</strong> the epilithic Caloplaca<br />

species tolerate dryness and live on relatively<br />

rain protected, <strong>of</strong>ten warm vertical surfaces and<br />

overhangs. Many species bargain for an<br />

“alternative habitat” a nutrient rich, eutrophic,<br />

rainy habitat (bird roosts, wall crowns etc.), as in<br />

the case <strong>of</strong> C. decipiens, C. saxicola and ( less<br />

pronounced) C. flavescens, in the case <strong>of</strong> others<br />

almost never abandoning the typical niche, such<br />

as C. cirrochroa, C. obliterans, C. proteus, C.<br />

chrysodeta and C. xantholyta.<br />

Silicate rock “favoring” C. arenaria, C.<br />

atr<strong>of</strong>lava, C. crenularia, the C. grimmiae living<br />

as a parasite on Candelariella vitellina as well as<br />

C. irrubescens, C. obliterans, C. scotoplaca, and<br />

C. subpallida: frequently occurring on relatively<br />

SiO2-poor, so called intermediary (“neutral”) or<br />

slightly calcareous or at best “mineral-rich”<br />

silicate rock. C. atr<strong>of</strong>lava lives also<br />

amphibiously on silicate boulders in cool brooks.<br />

C. cerina resides on base-rich barks, where<br />

closely related C. haematites, C. cerinella, C.<br />

cerinelloides, C. holocarpa, C. obscurella, and<br />

the threatened with extinction C. ferruginea, c.<br />

lobulata, C. hungarica and C. herbidella were<br />

earlier found in Heidelberg. C. lucifuga almost<br />

always occurs in rain sheltered bark crevices <strong>of</strong><br />

old oaks.<br />

C. tiroliensis (Black Forest), C. sinapisperma<br />

(E.g, Schwäbisch-Fränkischer Jura) and C.<br />

ammiospila (Sudeten) occur residing on moss<br />

and plant detritus in boulder fields sheltered cliff<br />

fissures distributed in the arctic-alpine, very<br />

isolated in the central European mediterranean<br />

region. Growing over mosses and plant detritus<br />

in the dry turfs and on calcareous rocks, C.<br />

cerina var. chloroleuca, is significantly more<br />

widely distributed, <strong>of</strong>ten in the montane zone and<br />

penetrating more deeply. The arctic-alpine moss<br />

dwelling C. jungermanniae, C. epiphyta and C.<br />

tetraspora, as well as C. aurea occurring on soil<br />

in rock fissures in the alpine regions <strong>of</strong> central<br />

and south Europe, are known in Germany only<br />

from the alpine (e.g. Allgäu).<br />

Many Caloplaca species are indigenous in<br />

south Europe and reach the north boundary <strong>of</strong> the<br />

area in central Europe, thus in southern central<br />

Europe C. aurantica, C. conversa, C. granulosa,<br />

C. irrubescens, C. ochracea, C. polycarpa, C.<br />

xantholyta and C. haematites in southern<br />

Scandinavia (in part only with isolated outposts)<br />

C. biatorina, C. chalybaea, C. cirrochroa, C.<br />

coronata, C. decipiens, C. dolomiticola, C.<br />

erythrocarpa, C. flavescens, C. lactea, C.<br />

teicholyta, C. variabilis, C. lobulata, C. lucifuga,<br />

C. obscurella and C. chrysophthalma. Perhaps<br />

thrusting further toward the north C. subpallida<br />

and C. cerinella. C obliterans occurs in north<br />

and central Europe. C. proteus and C. arnoldii<br />

have their high point in the calcareous mountains<br />

<strong>of</strong> southern central Europe and the mediterranean<br />

region. C. ferruginea and C. herbidella are<br />

predominantly westward distributed and limited<br />

in continental Europe to oceanic influenced<br />

mountain sites. C. hungarica is known from<br />

south Sweden, in the Alps and their environs. C.<br />

grimmiae is widely distributed in the drier<br />

regions. The area <strong>of</strong> C. cerina, c. citrina, C.<br />

flavovirescens, C. chrysodeta, C. crenularia, c.<br />

holocarpa and C. saxicola includes the greater<br />

part <strong>of</strong> Europe.<br />

Genus Characteristics and Determination<br />

Thallus very variable, crustose, mostly clearly<br />

developed, but also in the substrate, also rosette<br />

lobed (placoid), rarely squamulose (in the case <strong>of</strong><br />

exotic species even almost shrubby), yellow to<br />

red-orange, rarely whitish to dark gray or black,<br />

125

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