Revision of the Palaearctic species of the Coccinella ...

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136 KOVÁŘ: Coccinella transversoguttata species group and notes on other Coccinella Ventral surface moderately shiny, covered with dense, greyish-white, on distal part of tibiae pale ochraceous pubescence. Setae usually as long as 4-6 diameters, in middle of metasternum and of abdominal ventrites as 3-4 diameters, and on lateral parts of body at most as 6- 8 diameters of eye facet. Propleura horizontal, in anterior half widely longitudinally excavated, gradually impressed towards antero-lateral corners of prosternum, forming indistinct groove for reception of antennal tip. Surface obsoletely reticulate, shallowly punctate, punctures equal to eye facets or indistinct; small area close to antero-lateral corners of prosternum with fine dense oblique wrinkles. Epipleuron about 1.55 (1.47-1.69) times as wide as base of mesosternum, feebly sloping dorso-laterally, rather deeply and widely excavated in anterior 0.7. Surface obsoletely rugose, densely covered with shallow punctures of inequal size but usually larger than eye facets. Prosternum moderately convex in middle, its anterior margin rather deeply semi-elliptically emarginate. Basisternal lobes rather narrow, flattened, at the narrowest part distinctly more than twice as wide as prosternal process. Prosternal process narrow, only finely saddle-shaped behind midlength, apex semicircular. Prosternal carinae parallel, reaching anterior third of length of prosternum. Surface of prosternum finely transversely wrinkled, finely and rather densely punctate, punctures becoming obsolete postero-laterally. Mesosternal process at base 1.18 (1.06-1.29) times as wide as long in middle, rather strongly convex. Anterior margin moderately carinulate, straight or very shallowly emarginate in middle. Surface without wrinkles or indistinctly transversely wrinkled, sometimes obsoletely reticulate, coarsely and nearly regularly punctate. Punctures separated by their diameter, becoming denser towards base, 1.2-1.5 times as large as eye facets. Metasternum 3.97 (3.75-4.26) times as long as mesosternal process, rather strongly transversely convex, flattened posteromedially. Medial longitudinal sulcus complete, slightly impressed except for anterior portion. Precoxal bulge well developed, transverse. Surface finely transversely wrinkled and finely punctate. Wrinkles becoming finer and very dense antero-laterally. Punctures along medial sulcus inequal in size and distribution, 0.6-1.2 times as large as eye facets, somewhat deeply impressed, usually sparse, becoming somewhat denser and shallower to indistinct along lateral margins. Abdominal ventrites 1-4 in medial third distinctly transversely convex, sides flattened. Ventrite 1 only finely saddle-shaped medially, with surface rather coarsely densely punctate; punctures 1.2-1.5 times as large as eye facets, separated by 0.25-0.5 their diameter, becoming smaller and sparser postero-medially. Femoral line V-shaped, inner part strongly arcuate at basal half, then gradually straightened, shortly broken before meeting straight line before posterior margin of ventrite 1, and nearly crossing the outer, strongly oblique and nearly straight lateral line. Posterior margin of ventrite 5 in female straight, in male widely shallowly emarginate, that of ventrite 6 in female arcuate and distinctly acuminate, in male narrowly rather deeply arcuately emarginate. Moreover, ventrite 6 in male with shallow transversely oval impression in the middle. Legs long, slender, distal end of hind femora reaching a little over the outer margin of epipleuron. Hind tibia 7.44-8.02 times as longer as wide, outer margin feebly arcuate, inner one nearly straight. Tarsus moderately slender, tarsomere 3 reaching only a little over the midlength of the free part of tarsomere 2. Tarsal claw moderately slender, rather slightly curved dorsally, with sharp basal tooth situated submedially. Male genitalia. Tegmen moderately sized. Median lobe of aedeagus in larger basal portion subrectangular, in smaller apical one hastate, extended well beyond apices of parameres.
Acta Entomologica Musei Nationalis Pragae, 45, 2005 137 Hastate portion rhomboid and dilated at base, passing into narrow, tongue-shaped apex, distinctly shorter than half of hastate portion. Dorsal margin slightly sinuate with apex sharply pointed, feebly curved ventrally. Paramere thick, finger-shaped, only slightly curved at basal 0.4. Basal piece of aedeagus quadrangular, bearing large triangularly acuminate appendix. Trabes robust, gradually more strongly curved distally, nearly as long as basal piece and paramere combined. Sipho moderately long. Siphonal capsule robust with inner hook-shaped projection strong, curved dorsally, dorsal rib well developed, irregularly semicircular. Siphonal tube moderately slender, in basal third irregularly semicircularly rounded, forming rather sharp angle, apex of sipho only slightly dilated at the base. Preapical supporting sclerites parallel-sided dorsoventrally and narrowed laterally, bearing two inequal interconnected membranose swellings at each side, not extremely enlarged. Proximal pair of swellings subtriangular, without free portion, distal one oval, partly free. Terminal ampulla of sipho moderate in size, subtriangular. Female genitalia. Genital plate (hemiventrite 9) robust, about twice as long as wide, asymmetrically pear-shaped with rather thick, strongly curved basal portion. Stylus well developed, conical, bearing 4 setae. Spermatheca semicircular, nodulus short, much shorter than ramus, cornu moderately enlarged medially. Infundibulum about as long as spermatheca, with body cylindrical, equally enlarged at both ends, campaniform distal end rather short. Length. Males (n = 3) 4.87-5.28 mm; females (n = 3) 5.13-5.35 mm. KAPUR (1973) gives 5.4 mm. Variability. Black spots on elytra discrete (Fig. 83) or gradually confluent in a pattern of ½+S, 1+2+3 and 4+5 (Fig. 84) or of 1+2+3+S+½+S+3+2+1 (Fig. 85) to form a tricuspidate or anchor-like pattern in anterior half of elytra. Spots on posterior half of elytra invariably confluent. Distribution. Known material of C. marussii is extremely limited, coming only from the mountainous parts of northernmost Pakistan and Kashmir. Discussion. The species has been known only from the female holotype deposited in Museo Civico di Storia Naturale, Trieste, Italy. Although I have not studied the type, the identity of the examined material is doubtless given the locality. It is a characteristically coloured species with the tips of hind femora visible from above, and related to C. magnopunctata Rybakow, 1889, in having similar siphonal structures. The hastate portion of median lobe, however, much resembles that of C. transversoguttata. FÜRSCH (1981) also recorded C. marussii from Turkey but the material belongs to another, undescribed species; this new species is described below. Coccinella (Coccinella) hodeki sp. nov. (Figs. 21-33, 77, 86) Coccinella marussii: FÜRSCH (1981): 81 (misidentification). Type material. HOLOTYPE, : TURKEY: ERZURUM province, 15 km SW Erzurum, Palandöken Dağları [Mts.], Tekederesi village env., 39°48.8Ń, 041°09.0É – 39°47.6Ń, 041°09.7É, steppes and pasture near spring, tunnels of gopher, 1965-2230 m a.s.l., 26.-28.vi.2004, J. Hájek & J. Růžička lgt. Antenna, abdomen, male genitalia and tarsal claw of hind leg dissected.
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Revision of the Palaearctic species of the Coccinella ...

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