Revision of the Palaearctic species of the Coccinella ...

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148 KOVÁŘ: Coccinella transversoguttata species group and notes on other Coccinella trally, reaching over apex of paramere. Parameres strong, finger shaped, slightly curved in basal half. Basal piece quadrangular, somewhat wider than long, bearing subpentagonal basal appendix. Trabes robust, slightly curved distally, only a little shorter than basal piece and paramere combined. Sipho comparatively long, siphonal capsule robust, inner hook-shaped projection distinctly curved dorsally, roundly truncate at apex; dorsal rib strongly developed, semicircular. Siphonal tube in basal third widely rectangularly rounded, then straightened and slightly curved dorsally. Siphonal apex rather long, strikingly dilated at base (dilatation in lateral view elongate triangular dorsally), supporting sclerites parallel-sided dorso-ventrally (lateral view) and gradually narrowed anteriad (dorsal view), bearing three inequal swellings: two anterior ones as in C. marussii, and one round basal one armed with micro-prominences. Terminal ampulla of sipho comparatively small, subtriangular. Female genitalia. Genital plate (hemiventrite 9), robust, about twice as long as wide, asymmetrically pear-shaped with rather slender, moderately curved basal portion, broadly arcuately truncate at apex. Stylus well developed, subcylindrical, bearing four setae. Spermatheca C-shaped, rather slender, less curved medially than at both ends, cornu hardly enlarged in middle, nodulus shortly conical, ramus somewhat shorter than nodulus, quadrangular in outline. Infundibulum about as long as spermatheca, with body cylindrical and enlarged at both ends; basal portion very shortly cone-shaped, distal dilatation 1.5 times as long and asymmetrically campaniform. Length. Males (n = 3) 6.02-6.46 mm; females (n = 5) 6.32-6.91 mm. WEISE (1889), DO- BRZHANSKIY (1926) and MADER (1930 in 1926-1937) report 5-7 mm. Variability. No principal variability in the colour pattern was found; the elytral spots always remain isolated (Figs. 87-88). Distribution. China: Gansu, Quinghai, Sichuan and Xizang provinces; India: Kashmir; Mongolia (IABLOKOFF-KHNZORIAN 1979, 1982); Nepal: Manang district; Russia: Tuva. Discussion. Originally described as a variety of Coccinella undecimpunctata Linnaeus, 1758 by RYBAKOW (1889), who found unusually enlarged elytral black spots ½, 3 and 5 in an unknown number of specimens from localities ‘Dy-Tschu’ and ‘Amdo’ (in China, Quinghai province). WEISE (1889) recorded only the variety magnopunctata of C. undecimpunctata in provinces ‘Kan-ssu’ and ‘Sze-tschuan’ and based C. semenowi Weise, 1889 on other (numerous) material from ‘Kan-ssu’ and ‘Amdo’. His short description was based on differences in the position and size of some elytral spots between C. undecimpunctata and C. semenowi. SICARD (1892) added magnopunctata Ribakow [sic!] to the list of varieties of C. undecimpunctata. He was followed by GANGLBAUER (1899) and consequently by many students of the European Coccinellidae at the beginning of the 20 th century. JACOBSON (1915) regarded C. semenowi as a distinct species, giving its distributional data. DOBRZHANSKIY (1926) studied the types of C. magnopunctata and C. semenowi deposited in the Zoological Museum of the Academy of Sciences (St. Petersburg), shortly redescribed C. magnopunctata as a distinct species, synonymized C. semenowi with it, and figured the aedeagus. MADER (1930 in 1926- 1937, as C. magnoguttata) compared external characters of C. magnopunctata with those of C. undecimpunctata. IABLOKOFF-KHNZORIAN (1979, 1982) placed the species in Coccinella s. str. and designated the holotypes of C. magnopunctata and C semenowi.
Acta Entomologica Musei Nationalis Pragae, 45, 2005 149 The species undoubtedly belongs to C. transversoguttata species group. It seems to be a derived species within this group as suggested by the three preapical siphonal membranose swellings and the extremely enlarged hastate portion of the median lobe of aedeagus. It is evidently more closely related to C. marussii than to C. transversoguttata. Coccinella (Coccinella) transversoguttata transversoguttata Faldermann, 1835 (Figs. 56-74, 80-82, 89-92, 96) Coccinella transversoguttata Faldermann, 1835: 454; MULSANT (1850a): 117, partim; CROTCH (1874): 116, partim; WEISE (1879): 109; WEISE (1885): 28, partim; JACOBSON (1915): 982, partim (distribution); DOBRZHANSKIY (1924): 22 (female genitalia); DOBRZHANSKIY (1926): 21; MADER (1930 in 1926-1937): 150, partim (infrasubspecific variation); FÜRSCH (1981): 82; BIELAWSKI (1984): 417. Coccinella transversoguttata var. Sedakovi: WEISE (1889): 537 (misidentification). Coccinella geminopunctata Liu, 1962: 265. Coccinella (s. str.) geminopunctata: IABLOKOFF-KHNZORIAN (1979): 68 (as incertae sedis); IABLOKOFF-KHNZORIAN (1982): 362. Coccinella (s. str.) transversoguttata: IABLOKOFF-KHNZORIAN (1979): 67 (catalogue); IABLOKOFF-KHNZORIAN (1982): 364. Coccinella (s. str.) transversoguttata biinterrupta Iablokoff-Khnzorian, 1979: 67 (catalogue); IABLOKOFF-KHNZORIAN (1982): 365. Material examined. CHINA: GANSU, Xiahe (Labrang), 3300-3700 m a.s.l., 1.-15.vi.1989, 8 spec., V. Major lgt. (CVNC, NMPC); Shinlong Shan Mts., Yuzhong, 3200 m a.s.l., 6.-7.vii.1989, 3 spec., V. Major lgt. (CVNC); SHAANXI, Hua Shan Mts., 19.vi.1991, 1 spec., R. Dunda lgt. (NMPC); Qing Ling Shan Mts., road Baoji – Taibai vill., pass 35 km S of Baoji, 21.-23.vi.1998, 1 spec., O. Šafránek & M. Trýzna lgt. (CVNC); Qing Ling Shan Mts., 1500 m a.s.l., Hou Zen Zi county, 30 km SE of Taiban Shan Mt., 26.vi.1998, 1 spec., O. Šafránek & M. Trýzna lgt. (NMPC); SICHUAN, Sabde env., 3000 m a.s.l., 13.vii.1992, 1 spec., R. Dunda lgt. (NMPC); Kangding, vii.1992, 2 spec., R. Sauer lgt. (NMPC); Kangding, 3000 m a.s.l., 23.-27.vii.1995, 5 spec., J. Schneider lgt. (NMPC); Zhilong, vii.1992, 1 spec., R. Dunda lgt. (NMPC); XIZANG, ‘Tibet’, Quamdo – E env., (31°09Ń, 97°11É), ca 3500 m a.s.l., cultural steppe, 15.vii.1997, 5 spec., M. Trýzna & O. Šafránek lgt. (CVNC); ‘E Tibet’, Bomi env. (29°52Ń, 95°45É), mixed forest ca 3000 m a.s.l., 9-10.vii.1997, 5 spec., M. Trýzna & O. Šafránek lgt. (CVNC); YUNNAN, Weibaoshan Mts., 2800-3000 m a.s.l., (25°12Ń 100°24É), 29.-30.vi.1992, l spec., V. Kubáň lgt. (NMPC); Habashan Mts., S slope, 1 spec., V. Kubáň lgt. (NMPC); Cangshan Mts., 2500-3000 m a.s.l., (25°43Ń 100°06É), 24.vii.1992, 1 spec., V. Kubáň lgt. (NMPC); Yunlongshan Nat. Reserve, 50 km N of Lijiang, 24.-29.vi.1993, 5 spec., E. Jendek & O. Šauša lgt. (NMPC); Zhongdien env. 6.-8.viii.1995, 18 spec., J. Schneider lgt. (NMPC). KAZAKHSTAN: Tian Shan Mts., Chr. Bayulu, Dolon pass, 3200 m a.s.l., 16.-18.vii.1988, 7 spec., L. Chernyshev lgt. (NMPC). KYRGYZSTAN: Issyk-Kul Lake (Przevalsk env.), vii.1981, 25 spec., S. Pokorný lgt. (NMPC); Issyk Kul Lake, Grigoryevka, vii.1991, 2 spec., R. Sauer lgt. (NMPC); Tian Shan Mts., Teploklyuchevka, vii.1991, 5 spec. R. Sauer lgt. (NMPC). MONGOLIA: BULGAN, Dashinchilen Somon (256 km of Ulaanbaatar), 8.vi.1959, 1 spec., C. Purkyně lgt. (NMPC); KHENTEY, Somon Cencher, 2 spec., Mandal lgt. (NMPC); 20 km W of Bashiret vill., 1200 m a.s.l., (48°34.99 N, 110°03.92 E),
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Revision of the Palaearctic species of the Coccinella ...

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