A Checklist of the Beetles of Cuba with Data on Distributions and ...

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16 Table 4. Continued. Arthropods <strong>of</strong> Florida and Neighboring Land Areas, Vol. 18 Total Total End. Native Intro Predict. Erroneous ORDER COLEOPTERA Genera Species Species Species Species Species Species _________________________________________________________________________________________ 73. Lycidae 6 11 9 2 0 0 0 76. Lampyridae 8 37 29 8 0 0 0 78. Cantharidae 12 19 19 0 0 0 0 Bostrichiformia Bostrichoidea 81. Nosodendridae 1 1 1 0 0 0 0 82. Dermestidae 5 8 1 7 0 7 0 83. Bostrichidae 13 18 0 14 4 3 0 84. Anobiidae 17 23 12 10 1 3 0 Cucujiformia Lymexyloidea 85. Lymexylidae 1 1 0 1 0 0 0 Cleroidea 87. Trogossitidae 5 10 3 6 1 6 0 89. Cleridae 11 20 17 0 3 2 1 93. Melyridae 5 8 7 1 0 0 0 Cucujoidea 95. Sphindidae 2 2 2 0 0 0 0 97. Nitidulidae 15 32 1 28 3 3 0 98. Smicripidae 1 1 0 1 0 0 0 99. Monotomidae 2 4 3 1 0 0 0 103. Silvanidae 6 12 3 7 2 6 0 104. Passandridae 2 2 0 2 0 1 0 106. Laemophloeidae 6 7 2 5 0 3 0 108. Phalacridae 7 10 7 3 0 0 0 111. Cryptophagidae 1 1 0 1 0 0 0 113. Languriidae 3 4 1 3 0 0 0 114. Erotylidae 2 6 5 1 0 0 1 116. Biphyllidae 1 1 1 0 0 0 0 117. Bothrideridae 3 4 0 4 0 0 0 118. Cerylonidae 5 6 2 4 0 0 0 120. Discolomatidae 1 2 2 0 0 0 0 121. Endomychidae 1 1 0 1 0 0 0 122. Coccinellidae 31 43 19 22 2 0 1 123. Corylophidae 2 4 3 1 0 0 0 124. Latridiidae 5 6 3 3 0 1 0 Tenebrionoidea 125. Mycetophagidae 3 3 0 3 0 1 0 128. Ciidae 4 20 7 13 0 1 0 130. Melandryidae 1 1 1 0 0 0 0 131. Mordellidae 3 10 8 2 0 0 0 132. Rhipiphoridae 2 5 0 4 1 1 0 133. Colydiidae 13 17 3 14 0 1 0 134. Monommatidae 2 2 0 2 0 0 0 135. Zopheridae 2 3 1 2 0 0 0 140. Tenebrionidae 48 174 104 63 7 4 0 143. Oedemeridae 5 8 3 5 0 2 0 145. Meloidae 4 7 3 4 0 0 0 146. Mycteridae 3 5 3 2 0 0 0 151. Salpingidae 1 1 0 1 0 0 0 152. Anthicidae 8 20 4 13 3 2 1 153. Aderidae 2 2 0 2 0 0 0 154. Scraptiidae 2 2 2 0 0 0 0
Table 4. Continued. A <strong>Checklist</strong> <strong>of</strong> <strong>the</strong> <strong>Beetles</strong> <strong>of</strong> <strong>Cuba</strong> 17 Total Total End. Native Intro Predict. Erroneous ORDER COLEOPTERA Genera Species Species Species Species Species Species _________________________________________________________________________________________ Chrysomeloidea 155. Cerambycidae 42 230 144 80 6 9 9 156. Bruchidae 13 36 8 24 4 0 0 159. Chrysomelidae 85 374 306 68 0 9 6 Curculionoidea 161. Anthribidae 12 29 23 5 1 2 0 162. Belidae 1 1 1 0 0 0 0 163. Attelabidae 4 13 6 7 0 0 0 164. Brentidae 9 20 14 5 1 1 0 167. Curculionidae 134 385 232 131 22 12 0 __________________________________________________________________________________ Totals 954 2673 1495 1092 86 98 38 ORDER STREPSIPTERA Corioxenidae 1 1 0 1 0 1 0 Myrmecolacidae 1 1 0 1 0 0 0 Halictophagidae 1 1 0 1 0 0 0 __________________________________________________________________________________ Totals 3 1 0 3 0 1 0 Summary and origins <strong>of</strong> <strong>the</strong> beetle fauna <strong>of</strong> <strong>Cuba</strong> Summary <strong>of</strong> diversity The <strong>Cuba</strong>n beetle fauna is now known to contain 87 families, 954 genera, and 2,673 species (Table 4). This diversity can be compared <strong>with</strong> <strong>the</strong> known beetle fauna <strong>of</strong> o<strong>the</strong>r areas, such as <strong>the</strong> United States <strong>with</strong> 131 families; Canada <strong>with</strong> 129 families, 3526 genera, and 25,160 species; Mexico <strong>with</strong> 114 families; Brazil <strong>with</strong> 104 families, 4,351 genera, and 26,755 species; and Australia <strong>with</strong> 113 families (Costa 2000, Marske and Ivie 2003, Navarrete-Heredia and Fierros-López 2001). It is clear that additional families, genera, and species remain to be discovered in <strong>Cuba</strong>. There are some evident patterns in <strong>the</strong> <strong>Cuba</strong>n beetle fauna, as discussed below. An unbalanced fauna The family, subfamily, tribal and generic composition <strong>of</strong> <strong>the</strong> <strong>Cuba</strong>n beetle fauna is dissimilar in many ways from that found in <strong>the</strong> neighboring continental lands <strong>of</strong> <strong>the</strong> sou<strong>the</strong>rn United States, Mexico, and Central America. The most conspicuous feature is that many taxonomic groups are absent. This comparative difference is called a dis- harmonic or unbalanced fauna. It is a result <strong>of</strong> <strong>the</strong> long oceanic isolation <strong>of</strong> <strong>Cuba</strong> as an island. If <strong>the</strong>re had been free and obstructed movement <strong>of</strong> faunas between all <strong>the</strong>se regions and <strong>Cuba</strong> it would be expected that <strong>the</strong> higher-level taxonomic makeup <strong>of</strong> <strong>the</strong> beetles <strong>of</strong> <strong>Cuba</strong> would be similar to that <strong>of</strong> <strong>the</strong> neighboring continental lands. The actual dissimilarity is strong evidence that <strong>Cuba</strong> has not had a direct land connection to <strong>the</strong>se neighboring areas and, as a corollary, that <strong>Cuba</strong> has been long isolated from <strong>the</strong>se lands by a significant oceanic barrier to beetle dispersal and colonization. Islands which have always been so isolated are called oceanic islands. Table 5 is a list <strong>of</strong> beetle families which are known in continental warm-temperate Nearctic and tropical Neotropical lands adjacent to <strong>Cuba</strong> and which might ei<strong>the</strong>r be eventually discovered in <strong>Cuba</strong>, or whose absence helps to show <strong>the</strong> unbalanced nature <strong>of</strong> <strong>the</strong> <strong>Cuba</strong>n fauna. Generic and specific endemism and species swarms
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A Checklist of the Beetles of Cuba with Data on Distributions and ...

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