A Checklist of the Beetles of Cuba with Data on Distributions and ...
A Checklist of the Beetles of Cuba with Data on Distributions and ...
A Checklist of the Beetles of Cuba with Data on Distributions and ...
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10<br />
Arthropods <str<strong>on</strong>g>of</str<strong>on</strong>g> Florida <strong>and</strong> Neighboring L<strong>and</strong> Areas, Vol. 18<br />
<str<strong>on</strong>g>of</str<strong>on</strong>g> a multiplicity <str<strong>on</strong>g>of</str<strong>on</strong>g> users; from agr<strong>on</strong>omists, to ecologists,<br />
l<strong>and</strong>-use managers, tax<strong>on</strong>omists, quarantine<br />
entomologists, ecologists <strong>and</strong> biogeographers. Users<br />
can insert <str<strong>on</strong>g>the</str<strong>on</strong>g>ir own new data as <str<strong>on</strong>g>the</str<strong>on</strong>g>y come<br />
up<strong>on</strong> it.<br />
Organizati<strong>on</strong><br />
The core <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> work is a list <str<strong>on</strong>g>of</str<strong>on</strong>g> valid species<br />
names, syn<strong>on</strong>yms, <strong>and</strong> <str<strong>on</strong>g>the</str<strong>on</strong>g> more recent references<br />
about <str<strong>on</strong>g>the</str<strong>on</strong>g>se species in <str<strong>on</strong>g>the</str<strong>on</strong>g> scientific literature. The<br />
following explains <str<strong>on</strong>g>the</str<strong>on</strong>g> selecti<strong>on</strong> <strong>and</strong> organizati<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g><br />
<str<strong>on</strong>g>the</str<strong>on</strong>g> included data.<br />
Higher classificati<strong>on</strong>. The checklist generally<br />
follows <str<strong>on</strong>g>the</str<strong>on</strong>g> sequence <strong>and</strong> phylogenetic arrangement<br />
<str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> world list <str<strong>on</strong>g>of</str<strong>on</strong>g> beetle families, subfamilies,<br />
<strong>and</strong> tribes <str<strong>on</strong>g>of</str<strong>on</strong>g> Lawrence <strong>and</strong> Newt<strong>on</strong> (1995 <strong>and</strong><br />
at http:www. coleopsoc.org/). Arnett <strong>and</strong> Thomas<br />
(2000), <strong>and</strong> Arnett et al. (2002). <str<strong>on</strong>g>Data</str<strong>on</strong>g> <strong>on</strong> distributi<strong>on</strong><br />
<strong>and</strong> habits <str<strong>on</strong>g>of</str<strong>on</strong>g> beetles outside <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>Cuba</str<strong>on</strong>g> are generally<br />
from Lawrence (1982). The families are numbered<br />
so that finding <str<strong>on</strong>g>the</str<strong>on</strong>g>m in sequence is easier.<br />
Numbering from <str<strong>on</strong>g>the</str<strong>on</strong>g> world list is used because several<br />
families not now known from <str<strong>on</strong>g>Cuba</str<strong>on</strong>g> may be<br />
expected to be found in <str<strong>on</strong>g>the</str<strong>on</strong>g> future.<br />
Included species. The list c<strong>on</strong>tains all species<br />
<str<strong>on</strong>g>of</str<strong>on</strong>g> beetles which are known or reported to occur<br />
in <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>. If <str<strong>on</strong>g>the</str<strong>on</strong>g> known distribti<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g> a species<br />
suggests that it very probably occurs in <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>, but<br />
<str<strong>on</strong>g>the</str<strong>on</strong>g>re are seemingly no records for it, its name <strong>and</strong><br />
references are enclosed in paren<str<strong>on</strong>g>the</str<strong>on</strong>g>ses ( ). Some<br />
lists or catalogues <strong>on</strong>ly vaguely give distributi<strong>on</strong>s<br />
as “West Indies” or “Antilles.” If <str<strong>on</strong>g>the</str<strong>on</strong>g> total distributi<strong>on</strong><br />
suggests that this includes <str<strong>on</strong>g>Cuba</str<strong>on</strong>g> I have c<strong>on</strong>servatively<br />
included <str<strong>on</strong>g>the</str<strong>on</strong>g> species. The functi<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g><br />
such additi<strong>on</strong>s is to alert future workers to species<br />
which are named but not yet explicitly recorded<br />
from <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>. Such citati<strong>on</strong>s could undoubtedly be<br />
far more numerous than those I have included. If<br />
<str<strong>on</strong>g>the</str<strong>on</strong>g> species has been recorded in <str<strong>on</strong>g>the</str<strong>on</strong>g> past from <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>,<br />
but <str<strong>on</strong>g>the</str<strong>on</strong>g> applicati<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> name is now known to be<br />
an error, <str<strong>on</strong>g>the</str<strong>on</strong>g> name <strong>and</strong> references are enclosed in<br />
brackets [ ]. I ask that users <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> catalogue bring<br />
errors <str<strong>on</strong>g>of</str<strong>on</strong>g> omissi<strong>on</strong> or inclusi<strong>on</strong> to my attenti<strong>on</strong>.<br />
Citati<strong>on</strong>s. The author, year <str<strong>on</strong>g>of</str<strong>on</strong>g> publicati<strong>on</strong>, <strong>and</strong><br />
page <str<strong>on</strong>g>of</str<strong>on</strong>g> publicati<strong>on</strong> are given (when known) for each<br />
valid species. These data will help to locate <str<strong>on</strong>g>the</str<strong>on</strong>g> original<br />
references. When a species was originally placed<br />
in a different genus than now, I give this original<br />
genus name when it is known to me.<br />
Syn<strong>on</strong>omies. I list syn<strong>on</strong>ym names below <str<strong>on</strong>g>the</str<strong>on</strong>g><br />
valid name <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> species. I do not give complete<br />
lists <str<strong>on</strong>g>of</str<strong>on</strong>g> syn<strong>on</strong>yms for <str<strong>on</strong>g>the</str<strong>on</strong>g> species in this checklist.<br />
My intenti<strong>on</strong> is to give syn<strong>on</strong>omies <strong>on</strong>ly for names<br />
which have been applied to <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>n records.<br />
Catalogs. I do not cite catalogs or lists <str<strong>on</strong>g>of</str<strong>on</strong>g> names<br />
if no new tax<strong>on</strong>omic, distributi<strong>on</strong>al, or bi<strong>on</strong>omic<br />
informati<strong>on</strong> is given in <str<strong>on</strong>g>the</str<strong>on</strong>g>m. I do not give citati<strong>on</strong>s<br />
to <str<strong>on</strong>g>the</str<strong>on</strong>g> older primary literature which is included<br />
in Blackwelder (1944, 1957; reprinted in<br />
1982). The literature cited in Blackwelder is from<br />
1944 <strong>and</strong> earlier. To include all <str<strong>on</strong>g>the</str<strong>on</strong>g>se older references<br />
here would greatly increase <str<strong>on</strong>g>the</str<strong>on</strong>g> length <str<strong>on</strong>g>of</str<strong>on</strong>g> this<br />
listing.<br />
References. It is my intenti<strong>on</strong> to give references<br />
for “younger” literature <strong>on</strong>ly; i.e., that published<br />
since Blackwelder (1944, 1957) which has<br />
c<strong>on</strong>tributed data <strong>on</strong> <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>n species. I have also tried<br />
to include references to recent faunal summaries<br />
or revisi<strong>on</strong>s which may not c<strong>on</strong>tain direct informati<strong>on</strong><br />
<strong>on</strong> <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>n species but which may c<strong>on</strong>tain keys,<br />
descripti<strong>on</strong>s, or illustrati<strong>on</strong>s or o<str<strong>on</strong>g>the</str<strong>on</strong>g>r informati<strong>on</strong><br />
<strong>on</strong> species in <str<strong>on</strong>g>Cuba</str<strong>on</strong>g> or which may be expected in<br />
<str<strong>on</strong>g>Cuba</str<strong>on</strong>g>. Primary older literature not cited in<br />
Blackwelder can be found in appropriate issues <str<strong>on</strong>g>of</str<strong>on</strong>g><br />
“Zoological Record” or in <str<strong>on</strong>g>the</str<strong>on</strong>g> several catalogues<br />
cited. The relevant family chapters in Arnett <strong>and</strong><br />
Thomas (2000) <strong>and</strong> Arnett et al. (2002) are valuable<br />
references for all beetle families in <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>, <strong>and</strong><br />
are usually not cited for each family.<br />
Types. Type status <strong>and</strong> depository are given (if<br />
known). Older species usually do not have designated<br />
types. Co-types or syntypes from which lectotypes<br />
may be designated may be in collecti<strong>on</strong>s in<br />
Havana (<str<strong>on</strong>g>the</str<strong>on</strong>g> Gundlach collecti<strong>on</strong>), or in Paris<br />
(Chevrolat <strong>and</strong> Jaquelin du Val), or elsewhere (see<br />
table 3). Jacquelin du Val published new species<br />
names from specimens in both <str<strong>on</strong>g>the</str<strong>on</strong>g> Chevrolat <strong>and</strong><br />
Guérin-Méneville collecti<strong>on</strong>s, so those will probably<br />
have appropriate type material. Horn <strong>and</strong> Kahle<br />
(1935-1937) may be <str<strong>on</strong>g>of</str<strong>on</strong>g> help in locating historically<br />
important collecti<strong>on</strong>s <strong>and</strong> specimens.<br />
Type localities. I list published type localities (t.l.)<br />
<strong>on</strong>ly for endemic species. In many older works, no<br />
type localities were specified. In <str<strong>on</strong>g>the</str<strong>on</strong>g>se cases “<str<strong>on</strong>g>Cuba</str<strong>on</strong>g>”<br />
can be seen to serve as a type area. If this is <str<strong>on</strong>g>the</str<strong>on</strong>g><br />
case, I usually do not cite “<str<strong>on</strong>g>Cuba</str<strong>on</strong>g>.”<br />
Distributi<strong>on</strong>s. Distributi<strong>on</strong>s <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> species are<br />
presented in two ways.<br />
(A). They are first summarized as <str<strong>on</strong>g>Cuba</str<strong>on</strong>g> <strong>and</strong><br />
<str<strong>on</strong>g>the</str<strong>on</strong>g>n as a list <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> known major isl<strong>and</strong>s or countries<br />
outside <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>Cuba</str<strong>on</strong>g> from where <str<strong>on</strong>g>the</str<strong>on</strong>g> species is<br />
known. Distributi<strong>on</strong> <str<strong>on</strong>g>with</str<strong>on</strong>g>in <str<strong>on</strong>g>the</str<strong>on</strong>g> USA is by abbreviati<strong>on</strong>s<br />
<str<strong>on</strong>g>of</str<strong>on</strong>g> states at <str<strong>on</strong>g>the</str<strong>on</strong>g> edges <str<strong>on</strong>g>of</str<strong>on</strong>g> <str<strong>on</strong>g>the</str<strong>on</strong>g> range in <str<strong>on</strong>g>the</str<strong>on</strong>g><br />
USA. “Endemic” is used as a term to indicate that<br />
<str<strong>on</strong>g>the</str<strong>on</strong>g> species is naturally limited to <str<strong>on</strong>g>Cuba</str<strong>on</strong>g>. “Intro-