A Checklist of the Beetles of Cuba with Data on Distributions and ...

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104 Arthropods <strong>of</strong> Florida and Neighboring Land Areas, Vol. 18 Wooldridge, D.P. 1981. Three new species <strong>of</strong> Throscinus LeConte, <strong>with</strong> notes on o<strong>the</strong>r species (Coleoptera: Limnichidae, Cephalobyrrhinae). Coleop. Bull. 35: 217-221. 57. FAMILY HETEROCERIDAE The variegated mud-loving beetles Both adults and larvae construct tunnels in sand or mud at <strong>the</strong> edges <strong>of</strong> streams and ponds, where <strong>the</strong>y feed on algae or diatoms (see Pacheco 1964). SUBFAMILY HETEROCERINAE TRIBE HETEROCERINI Filiolus Pacheco 1964: 84 (endemic genus) F. guttatus (Kiesenwetter) 1851: 294 (Heterocerus); Chevrolat 1864c: 407; Gundlach 1891: 52; Pacheco 1964: 84 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Ci: Cayamas (t.l.) Collections. Washington (type) Neoheterocerus Pacheco 1964: 54 (key to species) N. angustatus (Chevrolat) 1864c: 407 (Heterocerus); Gundlach 1891: 52; Pacheco 1964: 62; Spangler 1981 Distribution. <strong>Cuba</strong>, Bahamas, Haiti, USA (IA-NY-FL) <strong>Cuba</strong> localities. Ca: California. CA: Baraguá. M: Cárdenas Collections. Gundlach no. 662 TRIBE TROPICINI Tropicus Pacheco 1964: 102; see Bameul 1995 T. bilineatus (Chevrolat) 1864c: 407 (Heterocerus); Gundlach 1891: 53; Pacheco 1964: 122; Ivie and Stribling 1984: 948; Mascagni 1989: 189 Distribution. <strong>Cuba</strong>, Costa Rica <strong>Cuba</strong> localities. PR: Soroa Collections. Gundlach no.688. Havana. Washington T. pusillus (Say) 1823: 200 (Heterocerus); Pacheco 1964: 106; Spangler 1981 Distribution. <strong>Cuba</strong>, Jamaica, Antigua, México, Honduras, Costa Rica, Panamá, USA (CA-PA-FL), Canada (ON) <strong>Cuba</strong> localities. Collections. INCERTAE SEDIS Heterocerus Fabricius 1792: 262 H. decimmaculatus Chevrolat 1864c: 407; Gundlach 1891: 53; Pacheco 1964: 122 (<strong>the</strong> identity <strong>of</strong> this species is unknown) Distribution. <strong>Cuba</strong>. Endemic? <strong>Cuba</strong> localities. M: Cárdenas Collections. Gundlach no. 710 REFERENCES Bameul, F. 1995. Un nouveau Tropicus Pacheco de la Guadeloupe (Coleoptera, Heteroceridae). Bull. Soc. Ent. France 100: 475-480. Ivie, M. A. and J. B. Stribling. 1984. Taxonomic and nomenclatorial notes on Caribbean Tropicus Pacheco (Coleoptera: Heteroceridae). Proc. Ent. Soc. Washington 86: 946-950. Mascagni, A. 1989. Tropicus alcicornis sp. n. dell’America meridionale. Boll. Soc. Ent. Ital. 120: 187-190. Pacheco, F. 1964. Sistematica, filogenia y distribucion de los Heteroceridos de America (Coleoptera: Heteroceridae). Monografias del Colegio de Post-Graduados: no. l, Chapingo, México. Pp. 155. 58. FAMILY PSEPHENIDAE The water penny beetles Larvae feed on algae on submerged rocks in streams, and adults occur on vegetation at stream edges, or at lights. SUBFAMILY PSEPHENINAE Pheneps Darlington 1936: 67 P. cubanus Darlington 1936: 68; Spangler 1973: 357; 1981: 167 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Ca: Rio El Manantiales. Ci: Soledad (t.l.). Gu: Rio Baracoa, at Tinta; Cabezades de Rio Judio. O<strong>the</strong>r records in Spangler 1981: 167 Collections. Cambridge (type) REFERENCES Spangler, P.J. 1973. Aquatic Coleoptera collected on <strong>the</strong> biospeleological expeditions. Resultats des expeditions biospéologique <strong>Cuba</strong>no- Roumaines à <strong>Cuba</strong> 1: 353-358. Spangler, P.J. 1981. Supplement to <strong>the</strong> aquatic and semiaquatic Coleoptera <strong>of</strong> <strong>Cuba</strong>. Resultats des expeditions biospéologique <strong>Cuba</strong>no- Roumaines à <strong>Cuba</strong> 3: 145-171.
59. FAMILY CNEOGLOSSIDAE This family is known only from Central and South America. Larvae and habits are unknown. 60. FAMILY PTILODACTYLIDAE The ptilodactylid beetles Adults may occur on vegetation and are attracted to lights. Larvae occur in leaf litter, rotted logs, and debris at stream edges. SUBFAMILY PTILODACTYLINAE Ptilodactyla Illiger 1807: 342 P. annulicornis Chevrolat 1870: 71; Gundlach 1891: 249 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. M: Cárdenas. Collections. Gundlach no. 619 P. carbonaria Chevrolat 1870: 72; Gundlach 1891: 250 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. SC: Santiago de <strong>Cuba</strong> Collections. Gundlach no. 1877 P. emarginata Chevrolat 1870: 71; Gundlach 1891: 248 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. M: Cienaga de Zapata Collections. Gundlach no. 1886 P. militaris Chevrolat 1870: 72; Gundlach 1891: 249 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Ha: Cojímar Collections. Gundlach no. 1308 P. ramicornis Chevrolat 1870: 70; Gundlach 1891: 247 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. M: Cienaga de Zapata Collections. Gundlach no. 1884 P. simplex Chevrolat 1870: 70; Gundlach 1891: 248 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. M: Cienaga de Zapata Collections. Gundlach no. 1885 P. turquinensis Zayas 1988: 32 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. SC: Pico Turquino (t.l.) Collections. Zayas (type) 61. FAMILY CHELONARIIDAE The turtle or chelonariid beetles Little is known <strong>of</strong> <strong>the</strong> biology <strong>of</strong> adults or larvae <strong>of</strong> this family. The genus Chelonarium is abundant in <strong>the</strong> Neotropics, Asia, and Australia. They live in moist to dry litter or under bark, and some may live in <strong>the</strong> refuse heaps <strong>of</strong> ants. A <strong>Checklist</strong> <strong>of</strong> <strong>the</strong> <strong>Beetles</strong> <strong>of</strong> <strong>Cuba</strong> 105 Chelonarium Fabricius 1801: 101 C. convexum Méquignon 1832: 246 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Collections. C. costatipenne Méquignon 1832: 244 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Collections. C. maculatum Méquignon 1834: 54 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Collections. C. problematicum Méquignon 1832: 244 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Collections. C. punctatum Fabricius 1801: 101; Jacquelin du Val 1856: 68; Chevrolat 1864: 618 Distribution. <strong>Cuba</strong>, Puerto Rico <strong>Cuba</strong> localities. Collections. C. sublaeve Méquignon 1833: 46 Distribution. <strong>Cuba</strong>. Endemic <strong>Cuba</strong> localities. Collections. 62. FAMILY EULICHADIDAE This family occurs in nor<strong>the</strong>rn California, China, India, and sou<strong>the</strong>ast Asia. The larvae live in streams. 63. FAMILY CALLIRHIPIDAE The family occurs in North, Central, and South America but seemingly not in <strong>the</strong> West Indies. SUPERFAMILY ELATEROIDEA 64. FAMILY ARTEMATOPODIDAE In <strong>the</strong> New World, this family occurs from North America, through México to South America. Larvae may feed on moss, lichens, or leaf litter. They are unknown in <strong>the</strong> Caribbean. 65. FAMILY BRACHYPSECTRIDAE These beetles live under bark in arid western North America, India, and Malaysia. A larva has been found in Miocene amber in <strong>the</strong> Dominican Republic, so <strong>the</strong>y might yet be found in <strong>the</strong> West Indies. 66. FAMILY CEROPHYTIDAE This family is known from Europe, eastern North America, and Central and South America.
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Arthropods of Flor
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Fernando de Zayas (1912-1983), Ento
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Superfamily Elateroidea 67. Eucnemi
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