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phylogenetic relationships and classification of didelphid marsupials ...

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174 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 322<br />

Character 115. dp3 trigonid: uni- or bicuspidRtricuspid.<br />

Cryptonanus unduaviensis:<br />

No unambiguous changes.<br />

Gracilinanus aceramarcae:<br />

No unambiguous changes.<br />

Gracilinanus microtarsus:<br />

Character 52. Postorbital process: absentRpresent.<br />

Gracilinanus agilis:<br />

No unambiguous changes.<br />

Gracilinanus emiliae:<br />

Character 34. Caudal scales: annularRintermediate.<br />

Lestodelphys halli:<br />

Character 2. Rhinarium: with 2 groovesRwith 1<br />

groove.<br />

Character 22. Pedal digits III <strong>and</strong> IV: IV longestRIII<br />

longest.<br />

Character 36. Tail tip: prehensileRnot prehensile.<br />

Character 57. Sagittal crest: absentRsmall.<br />

Character 81. Stapes: bicrurateRcolumelliform.<br />

Character 114. Height p2, p3: subequalRp3 taller.<br />

Character 120. m3 hypoconid position: labially<br />

salientRlingual to protoconid.<br />

Thylamys macrurus:<br />

No unambiguous changes.<br />

Thylamys pusillus:<br />

No unambiguous changes.<br />

Thylamys pallidior:<br />

Character 2. Rhinarium: with 2 groovesRwith 1 groove.<br />

Thylamys venustus:<br />

No unambiguous changes.<br />

Marmosops impavidus:<br />

No unambiguous changes.<br />

Marmosops noctivagus:<br />

No unambiguous changes.<br />

Marmosops incanus:<br />

Character 28. Mammary position: abdominal-inguinalRpectoral.<br />

Character 43. Nasal width: wider posteriorlyR<br />

uniformly narrow.<br />

Character 114. Height p2, p3: p2 tallerRsubequal.<br />

Marmosops parvidens:<br />

Character 105. Preprotocrista: stops at paraconeRmeets<br />

anterolabial cingulum.<br />

Marmosops pinheiroi:<br />

No unambiguous changes.<br />

Marmosa lepida:<br />

Character 94. Upper canine: simpleRposterior accessory<br />

cusp present.<br />

Marmosa murina:<br />

Character 13. Lateral carpal tubercles: present in<br />

adult malesRabsent.<br />

Marmosa demerarae:<br />

No unambiguous changes.<br />

Marmosa paraguayana:<br />

Character 32. Body pelage on tail: noneRsubequal.<br />

Marmosa regina:<br />

No unambiguous changes.<br />

Marmosa mexicana:<br />

Character 65. Palatine fenestrae: absentRpresent.<br />

Marmosa robinsoni:<br />

No unambiguous changes.<br />

Marmosa rubra:<br />

Character 52. Postorbital process: presentRabsent.<br />

Character 76. RTP-CTP contact: absent (gap)Rsutured.<br />

Tlacuatzin canescens:<br />

Character 66. Maxillary fenestrae: absentRpresent.<br />

Character 126. RES {a1+ a5, m1}: metacentricRacrocentric.<br />

Character 128. RES {a3+a10, m3}: metacentricRacrocentric.<br />

Monodelphis brevicaudata:<br />

No unambiguous changes.<br />

Monodelphis theresa:<br />

Character 9. Gular gl<strong>and</strong>: presentRabsent.<br />

Character 73. Secondary foramen ovale: absentR<br />

formed by lamina.<br />

Monodelphis emiliae:<br />

Character 76. RTP-CTP contact: absent (gap)Rsutured.<br />

Character 114. Height p2, p3: subequalRp3 taller.<br />

Monodelphis peruviana:<br />

Character 61. Parietal-mastoid contact: presentRabsent<br />

Hyladelphys kalinowskii:<br />

Character 96. Height P2, P3: subequalRP2 taller.<br />

Character 98. Upper milk premolar: large & molariformRvestigial.<br />

APPENDIX 6<br />

Key to the Didelphid Genera<br />

This key is intended for use on adult<br />

specimens (with completely erupted permanent<br />

dentitions). Juveniles (with dP3/<br />

dp3) <strong>and</strong> subadults (lacking dP3/dp3, but<br />

with incompletely erupted third premolars<br />

<strong>and</strong>/or fourth molars) may not key out<br />

accurately if they lack traits seen in<br />

conspecific adults (e.g., large size, postorbital<br />

processes, secondary foramina ovales,<br />

contact between left <strong>and</strong> right exoccipitals;<br />

Abdala et al., 2001). Although the following<br />

couplets are based, ins<strong>of</strong>ar as possible,<br />

on characters with unambiguously distinguishable<br />

alternative states, key results<br />

should always be checked by comparing<br />

specimens character-by-character against<br />

generic descriptions <strong>and</strong> cranial illustrations<br />

(figs. 37–54). Couplet 10, the only<br />

step in our key based on a single character,<br />

is obviously problematic if an atypical<br />

specimen is in h<strong>and</strong>. Users would be well<br />

advised, if an unsatisfactory result is<br />

obtained at any subsequent step, to follow<br />

the alternative sequence on the hypothesis<br />

that the species in question is polymorphic.<br />

External <strong>and</strong> craniodental measurements<br />

used to index size in this key are defined<br />

<strong>and</strong> abbreviated as follows: Head-<strong>and</strong>-

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