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Lithops - Au Cactus Francophone

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fig. 10 . lithops pseudotruncatella (bgr.) n.e. br.<br />

PROBLEM OF DETERMINING SPECIES<br />

OF THE GENUS<br />

The delimitation of species in this genus is complicated by the fact<br />

that the flowers of the various species show such little variation that they<br />

cannot he used to any extent for purposes of classification. It is true that the<br />

colour of the flower might be used, but whether this is a sufficient or safe<br />

character to go by I am not prepared to say. It is well known that flower<br />

colour frequently, depends on the kind of soil in which the plant grows. L.<br />

Helmuti is a very distinct species. As has already been stated it has a yellow<br />

flower and yet a white flowered form has been recorded. Is one justified in<br />

creating a new species on this evidence alone? Another objection to the use<br />

of flower colour in the delimitation of species is that these plants are usu<br />

ally found in the vegetative condition and it sometimes takes years before<br />

they flower. I have therefore discarded the flower as a systematic character<br />

and have preferred to use the upper surface of the leaf for purposes of classi<br />

fication. I do not want for a moment to say that the character chosen is<br />

absolutely satisfactory, but what character is ? After all, our whole conception<br />

of species is a subjective one. Some have a narrow idea of species and oth<br />

ers prefer to draw a wider ring to include many different plants in a single<br />

species. It sill be noticed that I have not mentioned—or at least only casu<br />

ally—the size of the plants. So far as I can judge there is only one species, L.<br />

Nelii which is characterised by its small size. The size of a succulent depends<br />

to a great extent on the amount of water available. If the water supply is low<br />

the plants are shrivelled and small, whereas after a rain these same plants are<br />

larger, and one would hardly believe them to be the same wilted plants seen<br />

21

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