Marine Resources Assessment for the Marianas Operating ... - SPREP
Marine Resources Assessment for the Marianas Operating ... - SPREP
Marine Resources Assessment for the Marianas Operating ... - SPREP
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AUGUST 2005 FINAL REPORT<br />
Acoustics and Hearing—Infrasonic, pattern sounds have been documented <strong>for</strong> fin whales (Watkins<br />
et al. 1987). Fin whales produce a variety of sounds with a frequency range up to 750 Hz. The long,<br />
patterned 15 to 30 Hz vocal sequence is most typically recorded; only males are known to produce<br />
<strong>the</strong>se (Croll et al. 2002). The most typical fin whale sound is a 20 Hz infrasonic pulse (actually an FM<br />
sweep from about 23 to 18 Hz) with durations of about 1 sec and can reach source levels of 184 to<br />
186 dB re 1 µPa-m (maximum up to 200) (Thomson and Richardson 1995; Charif et al. 2002). Croll et<br />
al. (2002) recently suggested that <strong>the</strong>se long, patterned vocalizations might function as male breeding<br />
displays, much like those that male humpback whales sing. While no data on hearing ability <strong>for</strong> this<br />
species are available, Ketten (1997) hypo<strong>the</strong>sized that mysticetes have acute infrasonic hearing.<br />
♦ Blue Whale (Balaenoptera musculus)<br />
Description—Blue whales are <strong>the</strong> largest living animals. Blue whale adults in <strong>the</strong> nor<strong>the</strong>rn<br />
hemisphere reach 22.9 to 28 m in length (Jefferson et al. 1993). The rostrum of a blue whale is broad<br />
and U-shaped, with a single prominent ridge down <strong>the</strong> center (Jefferson et al. 1993). The tiny dorsal<br />
fin is set far back on <strong>the</strong> body and appears well after <strong>the</strong> blowholes when <strong>the</strong> whale surfaces (Reeves<br />
et al. 2002). This species is blue-gray with light (or sometimes dark) mottling.<br />
Status—Blue whales are classified as endangered under <strong>the</strong> ESA and endangered by <strong>the</strong> IUCN Red<br />
List (Reeves et al. 2003). The blue whale was severely depleted by commercial whaling in <strong>the</strong><br />
twentieth century (NMFS 1998b). There is no designated critical habitat <strong>for</strong> this species in <strong>the</strong> North<br />
Pacific. The stock structure of blue whales in <strong>the</strong> North Pacific is uncertain (NMFS 1998b). There<br />
might be as many as five populations of blue whales in <strong>the</strong> North Pacific (NMFS 1998b).<br />
Habitat Preferences—Blue whales inhabit both coastal and oceanic waters in temperate and tropical<br />
areas (Yochem and Lea<strong>the</strong>rwood 1985). Feeding grounds have been identified in coastal upwelling<br />
zones off <strong>the</strong> coast of Cali<strong>for</strong>nia (Croll et al. 1998; Fiedler et al. 1998; Burtenshaw et al. 2004), Baja<br />
Cali<strong>for</strong>nia (Reilly and Thayer 1990), and off sou<strong>the</strong>rn Australia (Gill 2002). Blue whales also feed in<br />
cool, offshore, upwelling-modified waters in <strong>the</strong> eastern tropical and equatorial Pacific (Reilly and<br />
Thayer 1990; Palacios 1999). Moore et al. (2002) determined that blue whale call locations in <strong>the</strong><br />
western North Pacific were associated with relatively cold, productive waters and fronts.<br />
Distribution—Blue whales are distributed from <strong>the</strong> ice edges to <strong>the</strong> tropics in both hemispheres<br />
(Jefferson et al. 1993). Blue whales as a species are thought to summer in high latitudes and move<br />
into <strong>the</strong> subtropics and tropics during <strong>the</strong> winter (Yochem and Lea<strong>the</strong>rwood 1985). Data from both <strong>the</strong><br />
Pacific and Indian Oceans, however, indicate that some individuals may remain in low latitudes yearround,<br />
such as over <strong>the</strong> Costa Rican Dome (Wade and Friedrichsen 1979; Reilly and Thayer 1990).<br />
The productivity of <strong>the</strong> Costa Rican Dome may allow blue whales to feed during <strong>the</strong>ir winter<br />
calving/breeding season and not fast, like humpback whales (Mate et al. 1999).<br />
The range of <strong>the</strong> blue whale is known to encompass much of <strong>the</strong> North Pacific Ocean, from<br />
Kamchatka (Russia) to sou<strong>the</strong>rn Japan in <strong>the</strong> west, and from <strong>the</strong> Gulf of Alaska south to at least<br />
Costa Rica in <strong>the</strong> east (NMFS 1998). The only (presumably) reliable sighting report of this species in<br />
<strong>the</strong> central North Pacific was a sighting made from a scientific research vessel about 400 km<br />
nor<strong>the</strong>ast of Hawai’i in January 1964 (NMFS 1998). Blue whale call locations in <strong>the</strong> western North<br />
Pacific suggest that <strong>the</strong>re is an association between whale distribution and <strong>the</strong> Emperor Seamounts,<br />
<strong>the</strong> steep continental slope off <strong>the</strong> Kamchatka Peninsula, and <strong>the</strong> Aleutian Island chain (Moore et al.<br />
2002). Moore et al. (2002) noted a seasonal progression of call-location concentrations over <strong>the</strong><br />
seamounts in winter, <strong>the</strong> Kamchatka Peninsula and seamounts in spring, <strong>the</strong> Kamchatka Peninsula<br />
and waters between <strong>the</strong> seamounts and Aleutian Islands in summer, and <strong>the</strong> seamounts again in fall.<br />
Blue whale calls have been recorded off Midway and O’ahu (Northrop et al. 1971; Thompson and<br />
Friedl 1982; McDonald and Fox 1999); <strong>the</strong>se provide evidence of blue whales occurring within<br />
several hundred kilometers of <strong>the</strong>se islands (NMFS 1998b). The recordings made off O’ahu showed<br />
bimodal peaks throughout <strong>the</strong> year, suggesting that <strong>the</strong> animals were migrating into <strong>the</strong> area during<br />
summer and winter (Thompson and Friedl 1982; McDonald and Fox 1999). Evidence of blue whales<br />
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