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374 Sharks<br />

HEXANCHIDAE<br />

Cowsharks, sixgill, and sevengill sharks<br />

Diagnostic characters: Small to large sharks with slender to stout bodies, not eel-shaped. Head with 6 or<br />

7 pairs of long gill slits, the last in front of pectoral-fin origins, the first pair not connected across<br />

throat; no nictitating lower eyelids; snout short, acutely to bluntly pointed; nostrils without barbels or nasoral<br />

grooves; mouth very long and extending far behind the eyes; teeth of upper and lower jaws unlike at sides<br />

of mouth, uppers small, narrow, with a main cusp and often smaller cusplets, lowers very large, broad, compressed,<br />

and saw-like, with a series of cusps or large cusplets; short dermal gill rakers present on inner gill<br />

slits; spiracles present, small. A single dorsal fin, posterior to pelvic fins; anal fin present; caudal fin much<br />

less than half the total length, strongly asymmetrical, with a pronounced subterminal notch but the lower lobe<br />

very short. Caudal peduncle not depressed, without keels; no precaudal pits. Intestinal valve of spiral type.<br />

Colour: grey, blackish, or brown above, lighter below.<br />

6 or 7 gill slits<br />

a single<br />

dorsal fin<br />

Habitat, biology, and fisheries: These are moderately abundant, inshore to deep-water sharks, found in<br />

shallow bays down to the continental slopes and submarine canyons, near the bottom or well above it. They<br />

eat a wide variety of bony fishes, other sharks, batoid fishes, marine mammals, cephalopods, and crustaceans.<br />

They are taken in deep-water line fisheries for sharks off Cuba in the area and elsewhere, and are incidentally<br />

caught in trawls. Cow sharks are relatively unimportant but regular components of targeted shark<br />

fisheries and bycatches of other fisheries.They may snap during capture or when provoked.Divers encountering<br />

the larger species underwater have found them to be docile, and at least one species is the subject of<br />

ecotouristic dive trips.<br />

Similar families occurring in the area<br />

Chlamydoselachidae: the wide-ranging, deep-water frill shark<br />

(Chlamydoselachus anguineus) has a single dorsal fin and 6<br />

gill slits, but is distinguished by a nearly terminal mouth, first<br />

pair of gill slits connected across the throat, small, 3-cusped<br />

teeth in both jaws, and a long, almost eel-like body.<br />

No other sharks in the area have a single dorsal fin and 6 or 7<br />

gill slits.<br />

no precaudal<br />

pits<br />

teeth on left side (Hexanchus nakamurai) intestinal valve of spiral type<br />

nearly<br />

terminal<br />

mouth<br />

Chlamydoselachidae

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