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BOTANY Higher Secondary Second Year - Textbooks Online

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Difference between C 3 and C 4 photosynthetic pathways<br />

C 3 pathway C 4 pathway<br />

1. Photosynthesis occurs in meso - Photosynthesis occurs in meso -<br />

phyll cells. phyll and bundle sheath cells.<br />

2. The CO 2 molecule acceptor is The CO 2 acceptor molecule is<br />

RuBP. phosphoenol pyruvate.<br />

3. The first stable product is a 3C The first stable product is a 4C<br />

compound called 3 – PGA. compound called OAA.<br />

4. Photorespiration rate is high Photorespiration is negligible<br />

and leads to loss of fixed CO2. and it is almost absent. Hence,<br />

It decreases CO 2 fixation rate. it increases CO 2 fixation rate.<br />

5. Optimum temperature is 20 to Optimum temperature is 30 to<br />

25°C. 45°C.<br />

6. Examples of C 3 plants are rice, Examples of C 4 plants are maize,<br />

wheat and potato. sugarcane,Tribulus and<br />

Amaranthus<br />

5.2.7 Photorespiration or C 2 cycle<br />

In animals and bacteria, only one kind of respiration known as dark<br />

respiration occurs. This is not affected by the presence or absence of light.<br />

But in certain green plants, there are two distinct types of respiration –<br />

photorespiration and dark respiration. Respiration that occurs in<br />

photosynthetic tissues in the presence of light and results in increased rate<br />

of carbondioxide evolution is called photorespiration or light respiration.<br />

Photorespiration involves three organelles – chloroplasts, peroxisomes<br />

and mitochondria. Oxidation of RuBP in the presence of high oxygen<br />

is the first reaction of photorespiration. This reaction is catalysed by<br />

Rubisco* enzyme called carboxylase. It leads to the formation of 2C<br />

compound – phosphoglycolic acid and 3C compound PGA. When PGA<br />

is used up in the Calvin cycle, the phosphoglycolic acid is dephosphorylated<br />

to form glycolic acid in the chloroplasts.<br />

From the chloroplast, the glycolic acid diffuses into the peroxisome<br />

where it is oxidised to glyoxalic acid and hydrogen peroxide. In peroxisome<br />

from glyoxalic acid, glycine is formed.<br />

Note : * Rubisco = Ribulose bisphosphate carboxylase<br />

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