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BOTANY Higher Secondary Second Year - Textbooks Online

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3. Malic acid or aspartic acid formed in chloroplast of mesophyll<br />

cells is transferred to the chloroplasts of bundle sheath where it is<br />

decarboxylated to form CO 2 and pyruvic acid in the presence of<br />

NADP + specific malic enzyme.<br />

4. Now, second carboxylation occurs in chloroplasts of bundle sheath<br />

cells. Ribulose bisphosphate accepts CO 2 produced in step (3) in the<br />

presence of RuBP carboxylase and yields 3-phosphoglyceric acid. Some<br />

of the 3-phosphoglyceric acid molecules are utilised to produce sucrose<br />

and starch, while remaining PGA molecules are used for the regeneration<br />

of RuBP.<br />

5. The pyruvic acid produced in step (3) is transferred to the chloroplasts<br />

of mesophyll cells where it is phosphorylated to regenerate<br />

phosphoenolpyruvic acid . This reaction is catalysed by pyruvate kinase<br />

in the presence of Mg 2+ .<br />

Adenylate kinase<br />

The AMP is phosphorylated by ATP in the presence of adenylate<br />

kinase to form 2 molecules of ADP.<br />

C 4 plants are photosynthetically more efficient than C 3 plants, because<br />

the net requirement of ATP and NADPH 2 for the fixation of one molecule<br />

of CO 2 is considerably lower in C 4 plants than in C 3 plants.<br />

186

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