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BOTANY Higher Secondary Second Year - Textbooks Online

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cytochrome, DNA, RNA, manganese, etc. Chloroplasts are generally<br />

considerably larger than mitochondria.<br />

The space enclosed by the envelope is filled with matrix called stroma.<br />

In the stroma, many grana are embedded. In each granum, several disc<br />

shaped lamellae are found. These disc shaped structures are called<br />

thylakoids. They resemble a stack of coins. This structure is known<br />

granum. Generally a chloroplast contains 40 to 60 grana. The<br />

photosynthetic pigments are found in grana. The stroma contains circular<br />

DNA, RNA and enzymes for starch synthesis.<br />

5.2.3. Photochemical and biosynthetic phases<br />

The pigments involved in photosynthesis are called photosynthetic<br />

pigments. They are chlorophyll ‘a’, chlorophyll ‘b’, carotenoids,<br />

xanthophyll and phycobilins. Magnesium is an essential component for<br />

the formation of chlorophyll. Chlorophyll ‘a’ is a universal pigment present<br />

in the plants in which water is one of the raw materials for photosynthesis.<br />

Chlorophylls are highly efficient in absorbing solar energy and they are<br />

directly linked to photosynthetic electron transport. Photosynthetic pigments<br />

other than chlorophyll ‘a’ are generally called accessory pigments eg.<br />

chlorophyll ‘b’, carotenoids and xanthophyll, whereas chlorophyll ‘a’ is<br />

regarded as primary pigment.<br />

Photosynthetic pigments occur in the granum. They constitute the<br />

pigment system called photosystem. About 250 to 400 pigment molecules<br />

are present in a photosystem. Two types of photosystems are found in the<br />

granum. Photosystem I (PS I) has less accessory pigments and more<br />

chlorophyll ‘a’, while photosystem II (PS II) has more accessory pigments<br />

and less chlorophyll ‘a’. The primary function of photosystems is to trap<br />

light energy and converts it to chemical energy. The energy absorbed by<br />

accessory pigments is transferred to the chlorophyll ‘a’. The granal lamella<br />

where the photosynthetic pigments are aggregated to perform photosynthetic<br />

activities is called active centre.<br />

Mechanism of photosynthesis<br />

The overall reaction of photosynthesis can be written as follows.<br />

176

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