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BOTANY Higher Secondary Second Year - Textbooks Online

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points called chiasmata. Crossing over occurs between the nonsister<br />

chromatids of paired chromosomes in the region of chiasma. At each<br />

chiasma, the two nonsister chromatids break, exchange their segments<br />

and rejoin resulting the crossing over.<br />

5 M<br />

A ←⎯⎯⎯ ←⎯⎯⎯ 50 M ⎯⎯⎯→<br />

⎯⎯⎯→B←<br />

25 M →C←→D← 20 M →E<br />

Fig. 3.5 Illustration to explain the probability of crossing over<br />

Hence, out of four chromatids the two adjacent chromatids are<br />

recombinants and other two are original chromatids. Thus four types of<br />

gametes are obtained.<br />

Significance of crossing over<br />

Crossing over leads to the production of new combination of genes<br />

and provides basis for obtaining new varieties of plants.<br />

It plays an important role in the process of evolution.<br />

The crossing over frequency helps in the construction of genetic maps<br />

of the chromosomes.<br />

It gives us the evidence for linear arrangement of linked genes in a<br />

chromosome.<br />

Gene mapping<br />

Genes are arranged linearly in a chromosome. The point in a<br />

chromosome where the gene is located is called locus. The diagrammatic<br />

representation of location and arrangement of genes and relative distance<br />

between linked genes of a chromosome is called linkage or genetic map.<br />

The unit of genetic map is Morgan or centimorgan. When the<br />

percentage of crossing over between two linked genes is 1 per cent, then<br />

the map distance between the linked genes is one morgan.<br />

There is a greater probability of occurrence of crossing over, when the<br />

two genes are farther apart in a chromatid. The probability of crossing over<br />

between two genes is directly proportional to the distance between them.<br />

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