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BOTANY Higher Secondary Second Year - Textbooks Online

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chromosome has one centromere and the polycentric chromosome has<br />

many centromeres.<br />

The centromere contains a complex system of fibres called<br />

kinetochore. Each centromere has two kinetochores lined with<br />

chromosomal arms. The kinetochore is made up of protein fibres and<br />

microtubules which assist in the formation of spindles during mitosis<br />

and meiosis.<br />

All constrictions other than primary are called secondary<br />

constrictions. In a given set of chromosomes only one or two<br />

chromosomes have secondary constrictions. The nucleoli develop from<br />

secondary constrictions and such secondary constrictions are called<br />

nucleolar organisers.<br />

A satellite is a short chromosomal segment and separated from<br />

the main chromosome by a relatively elongated secondary constriction.<br />

A chromosome with a satellite is called SAT-chromosome.<br />

Chromatin is a viscous gelatinous substance that contains DNA,<br />

RNA, histone and non–histone proteins. H1, H2A, H2B, H3 and H4<br />

are the five types of histones found in the chromatin. The chromatin is<br />

formed by a series of repeated units called nucleosomes. Each<br />

nucleosome has a core of eight histone subunits.<br />

Telomere is the terminal part of chromosome. It offers stability<br />

to the chromosome. DNA of the telomere has specific sequence of<br />

nucleotides. Eukaryotic chromosome has DNA, RNA, histones, non–<br />

histone proteins and metallic ions like Ca +2 , Mg +2 , etc.,<br />

Types of chromosomes<br />

The chromosomes are classified into different types based on shape<br />

and position of the centromere. According to the position of centromere,<br />

Centromere<br />

Arm<br />

Telocentric Acrocentric Sub-metacentric Metacentric<br />

Fig. 3.2 Four morphogenic types of chromosomes<br />

111

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